The Project Gutenberg eBook of The book of antelopes, vol. 3 (of 4), by Philip Lutley Sclater

Title: The book of antelopes, vol. 3 (of 4)

Authors: Philip Lutley Sclater

Oldfield Thomas

Release Date: July 13, 2023 [eBook #71179]

Language: English

Credits: Peter Becker, Karin Spence and the Online Distributed Proofreading Team at https://www.pgdp.net (This file was produced from images generously made available by The Internet Archive)

[i]

THE
BOOK OF ANTELOPES.

BY

PHILIP LUTLEY SCLATER, M.A., Ph.D., F.R.S.,

SECRETARY TO THE ZOOLOGICAL SOCIETY OF LONDON,

AND

OLDFIELD THOMAS, F.Z.S., F.R.G.S.,

ASSISTANT IN THE ZOOLOGICAL DEPARTMENT OF THE BRITISH MUSEUM.

IN FOUR VOLUMES (1894–1900).

VOL. III.

LONDON:
R. H. PORTER, 7 PRINCES STREET, CAVENDISH SQUARE, W.
1897–1898.

[ii]

ALERE FLAMMAM.

PRINTED BY TAYLOR AND FRANCIS,
RED LION COURT, FLEET STREET.

VOL. III.

General Characters.—Size medium or small. Muzzle hairy. Anteorbital glands variable, large in some genera, absent in others. Tail generally short. Mammæ 2 (4 in Saiga).

Skull usually with supraorbital pits, lachrymal vacuities, and anteorbital fossæ. Molars tall and narrow.

Horns present in the male only, except in Antidorcas and in most of the species of Gazella.

The greater part of this subfamily consists of the Gazelles and their allies, the Saiga, Chiru, Springbuck, Gerenuk, and Dibatag; and with these, by common consent, are included the Black-buck, the typical form of the whole group of Antelopes, and the Pallahs. We have also thought that the anomalous little Antelope known as the Beira would best be placed in this subfamily, near the Gazelles, in spite of the superficial resemblance it bears to certain members of the subfamily Neotraginæ.

[2]

The present subfamily consists therefore of nine genera, which may be arranged as follows:—

• A. Horns spirally twisted  1. Antilope.
• B. Horns curved or straight, not twisted.
• a. False hoofs absent  2. Æpyceros.
• b. False hoofs present.
• a¹. Horns medium or long, curved.
• a². Muzzle swollen or elongated.
• a³. Horns medium, lyrate, whitish  3. Saiga.
• b³. Horns long, slightly curved, black  4. Pantholops.
• b². Muzzle slender, normal.
• a³. Neck normal.
• a⁴. Horns convex forwards for three-fourths their length.
• a⁵. Back with a central white streak. Lower premolars 2.  5. Antidorcas.
• b⁵. Back normal. Lower premolars 3  6. Gazella.
• b⁴. Horns concave forwards, except just at their base.  7. Ammodorcas.
• b³. Neck much elongated. Horns as in Gazella.  8. Lithocranius.
• b¹. Horns short, quite straight  9. Dorcotragus.

Size medium. Muzzle hairy. A large anteorbital gland present. Tail short, compressed. Mammæ 2. Accessory hoofs present. Glands in all the feet and in the groin.

Skull with deep pits between the orbits, very small or no lachrymal vacuities, and large anteorbital fossæ. Molars tall and narrow.

Horns long, placed close together, widely divergent, cylindrical, spirally twisted, closely ringed throughout. Female normally hornless.

One species only is known.

THE BOOK OF ANTELOPES, PL. XLVII.

J. Smit, del. & lith.

Hanhart imp.

The Black-buck

ANTILOPE CERVICAPRA.

Published by R. H. Porter.

[5]

76. THE BLACK-BUCK.
ANTILOPE CERVICAPRA (Linn.).
[PLATE XLVII.]

[7]

Height of male at withers about 30 inches. General colour in the same sex brown, gradually darkening with age to deep shining black. Muzzle and chin, an area round the eyes, and the whole of ears white. Back of neck, especially in the black individuals, yellowish. Upper part of flanks with an indistinct narrow whitish line running along them, most conspicuous in the young. Chest, belly, and inner sides of limbs pure white; outer sides of the latter brown. Tail short, its upperside fawn or brown, beneath white; its end with an indistinct blackish tuft.

Female brownish fawn wherever the male is black, and with the colour-contrasts nowhere so conspicuous. Back of ears and nape of neck also fawn. Horns absent, except in abnormal cases (see p. 14).

Skull as described above. The dimensions of a skull of a male are:—Basal length 8·3 inches, greatest breadth 4·0, muzzle to orbit 4·9.

The Indian Antelope or Black-buck, as the male is universally called by sportsmen, is usually associated with the Gazelles, and we retain it in this position, although it deviates from all the other members of the subfamily in having its horns spirally twisted somewhat after the manner of the Tragelaphinæ. It likewise differs from the rest of the group as regards the strong contrast of colour between the sexes, although this is of course a comparatively trifling character.

This Antelope, although strictly confined to India south of the Himalayas, has been more or less known in Europe for a long period, probably since the invasion of India by Alexander the Great. It has been even conjectured that the twisted horn of the fabled Unicorn of mediæval writers may have been originally based upon single horns of the present animal, though other authorities are inclined to refer the Unicorn’s horn to the Narwhal. This, however, is rather an antiquarian than a zoological question.

In the two last and most complete editions of the ‘Systema Naturæ’ Linnæus based his Capra cervicapra upon the descriptions of several of his predecessors (Gesner, Aldrovandus, Ray, and Brisson), which certainly refer to the present species, and we may therefore safely adopt cervicapra for it as its specific term. The name bezoartica of Linnæus, which has been employed[8] in its place by some authorities, refers to quite a different animal, probably to one of the wild goats, but certainly not to the Indian Antelope.

As regards the generic appellation of the present animal, we have already explained our reasons for following the general practice of the best modern authors in considering the Capra cervicapra of Linnæus to be the type of the genus Antilope, although Pallas, who founded the genus, did not give it precedence in his list of species. But the fact is that Pallas in his day never realized the importance attached in modern times to the exact designation of the types of genera, and had probably no intentions in the matter. The correct scientific name of the Black-buck is therefore, in our opinion, Antilope cervicapra.

The authors immediately subsequent to Linnæus, whose numerous references we quote in our synonymy, added little or nothing to our knowledge of the Indian Antelope. Shaw and other writers of the same date continued the story (which originally arose from its being confounded with the Addax) of its being met with in Africa as well as India—a fallacy which appears to have been first exposed by Lichtenstein in his excellent article on the genus Antilope, published in 1814. But accurate information on this Antelope and its exact range and habits was only obtained when the fauna of the Indian Peninsula came to be investigated by those whom the increase of English influence caused to be resident in that country.

After General Hardwicke, the late Sir Walter Elliot was among the first of the British residents in India who turned his special attention to the zoology of British India. In 1839 he published an excellent article upon the mammals of the Southern Mahratta country. Here, he tells us, the Indian Antelope “frequents the plains in herds of from twenty to thirty, each of which contains only one buck of mature age, the others being young ones.” In some cases the herds are so large that one buck has fifty or sixty does in its company, while the younger bucks, driven away by the old ones, wander about in separate herds, which sometimes contain as many as thirty individuals of different ages.

Jerdon, in his ‘Mammals of India,’ published in 1867, following Gray, calls the Indian Antelope Antilope bezoartica, but gives us a good account of it. It is found, he says, throughout India in suitable localities, but is not met with elsewhere. “It is rare in Bengal, a few only extending into Purneah and Dinagepore, north of the Ganges; and it does not occur in the richly[9] wooded Malabar coast. It is abundant in the Deccan, in parts of the Doab between the Jumna and Ganges, also in Hurriana, Rajpootana, and the neighbouring districts. It is found in the Punjab, but does not cross the Indus.”

McMaster, in his ‘Notes on Jerdon’s Mammals,’ and Sterndale, in his ‘Natural History of the Mammalia of India and Ceylon,’ besides numerous other writers in the ‘Bengal Sporting Magazine’ and other periodicals, have published good field-notes upon the Black-buck, which is perhaps the most favourite object of pursuit of the sportsman in the plains of India. But one of the best summaries of all these observations is that put together by General Kinloch in his excellent work on ‘Large Game Shooting in Thibet, the Himalayas, Northern and Central India,’ from the third edition of which, published in 1892, we venture to quote the following extracts:—

The chase of the Black-buck by the Cheetah (Cynælurus jubatus) is a favourite sport of the native Princes and Nobles of India. General Kinloch, in the work we have just quoted, describes one of these chases, in which he took part, as follows:—

Excellent accounts of the distribution and habits of the Indian Antelope have also been lately published in Dr. Stanford’s ‘Mammals of British India,’ and in the second volume of ‘Big Game Shooting’ in the Badminton Library. In the latter we find described the following curious method of getting within close range of the Black-buck as practised in Central India:—

The Indian Antelope bears captivity easily, and specimens of it are to be seen in all the Zoological Gardens of Europe, in some of which it has bred and multiplied very successfully. In other places it has not done so well, apparently requiring a light soil and a considerable amount of protection from the inclemencies of a northern climate.

In the celebrated Menagerie at Knowsley fifty years ago this Antelope is stated by Gray to have bred but once at the time he was writing of it (1846). But shortly afterwards the herd of this animal in Lord Derby’s possession appears to have increased very rapidly. When the Menagerie was dispersed by auction after the Earl’s death in 1851 we find that four males and four females of this Antelope were entered in the sale-list, all described as having been bred at Knowsley. These passed into the possession of the late Viscount Hill, of Hawkstone, who at that epoch shared Lord Derby’s tastes in his love for keeping living animals.

So far as we can tell from an inspection of the Zoological Society’s records, the first specimens of the Black-buck received by the Society were brought home by Col. Sykes (a well-known authority on Indian zoology) from Bombay in 1831. In the ‘Proceedings’ of the Society for 1836 Mr. E. T. Bennett, then Secretary of the Society, published some interesting remarks on this herd, especially referring to the vexed question of the use of the lachrymal sinus in Antelopes, which, from consideration of the relative development of it in the several specimens then in the Society’s Gardens, he showed was in all probability subservient to sexual purposes.

As will be seen by reference to the nine published editions of the ‘Lists of Animals in the Society’s Collection,’ numerous specimens of the Black-buck have been acquired by the Society since that date, but, probably on account of the small free space assigned to them, little or no success has been met with in breeding this beautiful species in the Regent’s Park. On the other hand, at the Jardin d’Acclimatation at Paris and in other places under a climate more genial than our own, where large paddocks can be assigned to it, the Black-buck frequently reproduces in captivity and flourishes exceedingly.

No figures of the Black-buck having been drawn under the late Sir Victor Brooke’s directions, our illustrations of this beautiful Antelope (Plate XLVII.)[13] have been taken by Mr. Smit from two mounted specimens in the British Museum of Natural History, the male of which, from Gwalior, was presented by Mr. C. Maries, of the Gwalior Museum.

Fig. 45.

Horns of Black-buck, ♂. (In the Collection of Mr. A. O. Hume.)

Fig. 46.

Abnormal horns of female Indian Antelope.
(In the collection of Mr. A. O. Hume.)

The National Collection possesses other examples of this Antelope, besides a fine series of heads and horns, mostly from Rajpootana and the Punjáb, belonging to the splendid collection of these objects presented to the British Museum by Mr. A. O. Hume, C.B. In order to show the large dimensions to which the horns of the adult male Black-buck attain in Rajpootana and Hurriana we give a drawing (fig. 45) of a beautiful pair still in the possession of Mr. Hume, which attained a length of no less than 28¼ inches measured in a straight line. On referring to the long list of the dimensions of the horns of[14] this Antelope published in Mr. Rowland Ward’s ‘Records of Big Game,’ it will be found that only one pair of greater length than the horns which we now figure have been hitherto recorded. We also give (fig. 46) a figure of the skull and abnormal horns of a female of this Antelope in Mr. Hume’s Collection.

August, 1897.

Size large. No anteorbital glands. Tail fairly long. False hoofs absent. Hind feet with glandular tufts of hair placed shortly above the hoofs.

Skull without supraorbital pits or anteorbital fossæ; lachrymal vacuities small.

Horns of male medium or rather long, broadly lyrate, half-ringed, slightly compressed. Female hornless.

Of this genus we are at present prepared to recognize only two species—the Common Pallah of Southern and Eastern Africa (Æ. melampus) and that of Angola (Æ. petersi). The latter may be readily distinguished from the ordinary form by having a prominent blackish mark running down the upper surface of the muzzle.

THE BOOK OF ANTELOPES, PL. XLVIII

Wolf del, J. Smit lith

Hanhart imp.

The Pallah

ÆPYCEROS MELAMPUS.

Published by R. H. Porter.

[17]

77. THE PALLAH.
ÆPYCEROS MELAMPUS (Licht.).
[PLATE XLVIII.]

Size comparatively large; height at the withers about 36 inches. General colour bright reddish brown, paler along the lower part of the sides. In southern specimens the colour is rather duller and browner than in northern ones, but the difference is very slight. Head dark fawn; a mark over the anterior half of the eye, chin, interramia, and upper part of throat white. Belly pure white. In front of the eye, on the side of the face, there is generally, though not always, in southern specimens an indistinct darker patch, but this is never present in northern ones. A black patch occasionally present on the crown. Ears of medium length, their outer sides fawn, with the terminal third black. Limbs like the back, a lighter ring round the[19] pasterns just above the hoofs; a pair of prominent black tufts of longer hairs on the distal extremity of the hind cannon-bones (whence the name of the species). Tail fairly long, its upperside with a narrow black line along it, extending more or less on to the back, its sides fawn basally, white terminally.

Skull as above described. The dimensions of a male skull are:—Basal length 10·3 inches, greatest breadth 4·4, muzzle to orbit 6·1.

The horns are particularly graceful, lyrate, convex forwards below, concave above, evenly spreading. In length, in the south, good specimens may attain about 18 or 20 inches in a direct line, and in the north more, up to about 21 or 22 inches, the largest recorded being 23. But in the intermediate districts, Nyasa, Zambesia, and Gazaland, they are much shorter, fully adult horns being often only 14 inches in length.

Female similar to the male, but without horns.

The first account of the Pallah seems to have appeared in one of the early numbers of a work called Daniell’s ‘Illustrations of African Animated Nature and Scenery,’ published in London in 1812. The author of the letterpress, however, did not give it a scientific name, believing that it might be the “Kob” of Buffon, or an allied species. At about the same date Prof. Lichtenstein, who had met with this animal during his journeyings in Southern Africa from 1803 to 1806, published a description and figure of it in his ‘Reise nach südlichen Afrika’ under the name Antilope melampus. This description, with additional particulars, was repeated in the same author’s classical monograph of the genus Antilope, published in 1814, and his name, taken from the black tufts of short hair at the back of the hind legs just above the foot (which are clearly shown in our figures), has been employed, almost universally, for this species by subsequent writers. Lichtenstein met with his specimens near Klip Fontein in Namaqualand, where it was found to occur in small herds of five or six individuals. In 1812 the celebrated African traveller Burchell likewise met with the Pallah in Bechuanaland, and secured the first specimens which arrived at the British Museum.

Little more was added to our knowledge of this beautiful Antelope until[20] the publication of Harris’s ‘Portraits of the Game and Wild Animals of Southern Africa’ in 1840. Harris devotes his fifteenth plate mainly to the illustration of the “rare and graceful Pallah,” which he states “first gladdens the sight of the traveller in Southern Africa upon the elevated districts north of Latakoo.” Here in the wooded slopes and valleys that environ the mountain-ranges of Kurrichane and Cashan it was met with in families of from twelve to twenty individuals of both sexes.

Harris, with all his experience, could recall to his memory “few objects more picturesque than the graceful figures of a wandering herd of these Antelopes dancing and bounding through the thousand stems of the acacia-groves in all the poetry of motion.” To these wooded districts Harris considered the Pallah to be restricted, not a single specimen having been observed in the open country. The flesh of the Pallah he characterizes as “tender and palatable,” although “rather dry,” like that of most Antelopes.

In these days, however, as we are informed by Messrs. Nicolls and Eglington, it is only on rare occasions that the Pallah is met with in the Bechuanaland Protectorate, and its present distribution is described by them as follows:—“A few herds still linger in the Transvaal along the Crocodile River. Almost exterminated in the regions through which the north-west tributaries of that river flow, it is only when the Zambesi is reached that the Palla is again to be frequently met with in any number. On the Chobe River it is still fairly common, being unknown on the Botletle, but it is only after passing the shores of Lake ’Ngami, and reaching the densely wooded banks of the Tonke, that the species again makes its appearance in a westerly direction. In those parts of Mashonaland and Matabeleland where it is not subject to continual persecution it is still fairly numerous. The Palla is highly gregarious, and frequents the thick, forest-clad banks of rivers, from which it never strays, except after periods of heavy rains, and then only when the pans and vleys (which are always dry during the greater portion of the year) are for a time filled with water. In remote parts, not very much frequented by man, the herds often exceed a hundred in number. Where not continually disturbed, this Antelope, so elegant and graceful of motion, is not by any means shy when approached, generally running but a short distance, and then standing and looking back again, a habit which easily permits of its being stalked.”

In the Transvaal, Mr. Barber kindly informs us, the Pallah was plentiful[21] in the Waterburg and Lydenburg districts up to 1880. Now, however, it has been driven away many miles east, into the valleys that intersect the Lebombo range.

On the north-west of the Cape Colony the Pallah, as we shall see presently, is represented by a nearly allied, though probably distinct, form. But on the eastern side of Africa the Pallah has a wide range, and extends north certainly into British East Africa, and probably still farther into Kordofan. We will endeavour to trace its range throughout this wide area.

Mr. Selous found the Pallah on the tributaries of the Limpopo, and thence northwards on the banks of every river and stream which he has explored in Matabeleland and Mashonaland. The Impalas of the Limpopo he considers to be larger than those of the Chobe.

Peters records the Pallah as met with in the mountainous parts of the Portuguese province of Mozambique, near Tette, Chidima, and Sena, and gives its native names as here ‘Psuara’ or ‘Suara.’ Passing into British Central Africa we find this Antelope recorded by Mr. Crawshay as not common anywhere in Nyasaland, but where met with, as a rule, found in even larger numbers than the Waterbuck. Mr. Crawshay has seen it in companies of one hundred or more, and gives a number of localities around Lake Nyasa in which he has come across herds of it. No Antelope, Mr. Crawshay tells us, can compare with the Pallah in fleetness of foot, and certainly “no other can display such wonderful leaping powers. They go off like the proverbial arrow from the bow, and with most beautiful gliding bounds, cover the ground without apparently the least effort.” In Northern Nyasaland, Mr. J. B. Yule tells us, the Pallah is found only along the stony ridges between Deep Bay and Karonga.

In the highlands of Zomba and the adjacent districts of Nyasaland a local race of the Common Pallah is found, distinguished by its slenderer skull and much shorter horns; but as regards the colour of its fur it is precisely similar to the South-African form. Thomas was at one time of opinion that this highland form should constitute a separate subspecies, and proposed to name it after its discoverer, Sir Harry Johnston, who has done so much in investigating the fauna of British Central Africa, Æpyceros melampus johnstoni. Thomas, however, since the examination of further specimens is not disposed to insist upon the necessity of recognizing this subspecies as distinct.

In the low, dry, thicket-covered hills to the north of Lake Mweru both[22] Mr. Crawshay and Mr. Sharpe have obtained specimens of this Antelope, which, according to the latter, is often confounded by the natives with the Lechee and Vardon’s Antelope under the common name “msala.”

In German East Africa, according to Dr. Matschie’s excellent Handbook, the Pallah occurs in many localities all over the country. At first misled by the association of the horns of a Lesser Koodoo and a skin of a Pallah, Dr. Matschie proposed to found a new species of Koodoo upon this animal, and to call it Strepsiceros suara. Afterwards recognizing his mistake he proposed to retain the term suara for the East-African Pallah, and to separate it specifically from the South-African animal as Æpyceros suara, on the ground of certain small discrepancies in colour. But after examining many specimens of the Pallah from East Africa we have come to the conclusion that the differences pointed out by Dr. Matschie are not confined to individuals from the same locality, and we cannot therefore regard Æ. suara as a distinct species.

The late Mr. F. Holmwood, formerly H.B.M. Consul-General at Zanzibar, wrote to us, “I have met with the Pallah in the countries of Usagara and Uzeguha, about 150 miles straight inland from Zanzibar, where they were very plentiful. The country has an elevation of 500 feet and is well watered. The Pallah go in troops of from 15 to 120. I once saw a pack of wild dogs hunt and run down one of these Antelopes which they first separated from a large herd.”

In British East Africa the Pallah is well known, and has been obtained by all the great sportsmen that have visited that territory. Mr. H. C. V. Hunter, in his appendix to Sir John Willoughby’s ‘East Africa,’ speaks of this Antelope as “common everywhere in thin bush and on the plains.” Dr. Abbott, as recorded by Mr. True, sent to Washington a good series of specimens obtained in 1889 from Taveta and Mount Kilimanjaro, where it had likewise been met with by Sir Harry Johnston during the Kilimanjaro Expedition of 1884. Mr. F. J. Jackson, in his notes on Antelopes published in the first volume of ‘Big Game Shooting’ of the Badminton Library, tells us that the Pallah is not met with in the coast-district of British East Africa. “But it occurs in small herds about 60 miles inland, and is plentiful at Adda and in the Teita country, and is found as far north as Turkwel in suitable localities, that is, in park-like open bush and thinly-wooded country, not far from water.” “The best heads,” Mr. Jackson says, “are obtained between[23] Lakes Navaisha and Baringo, particularly in the vicinity of the small salt-lake Elmatita, where these beautiful beasts inhabit the open woods of juniper-trees.” In his paper on the Antelopes of the Mau district, recently read before the Zoological Society, Mr. Jackson likewise speaks of this Antelope, and again mentions the large size of the horns of the bucks in that part of British East Africa, which he gives as 22 and 23 inches from base to tip.

Fig. 47.

Head of Pallah, ♂, front view.

Whether the Pallah ranges further north than British East Africa and the neighbouring district of Turkwel is perhaps not quite certain, though it may possibly be the case. Our only authority on the subject is Heuglin, who states that the Pallah occurs on the White Nile at Scherk-el-Akaba, and is “very common” on the Djur River, where it is known by the Arabic name of ‘Om-gàba,’ or ‘Om-sàba.’ But Heuglin’s observations on this point, so[24] far as we know, have not been confirmed, and we have never seen specimens from this locality.

On the whole, therefore, we consider Æpyceros melampus to be a wide-ranging species, extending from Bechuanaland in the south throughout the eastern side of Africa to British East Africa on the north, and perhaps reaching even to the White Nile. But over all these districts there is a certain amount of variety amongst the specimens, and we are not, therefore, at present inclined to recognize, even as subspecies, what have been designated as suara, johnstoni, and holubi, although future researches may lead us to a different conclusion.

So far as we know, the Pallah has been brought to Europe alive on two occasions only, and in both instances the animals were imported by Mr. C. Reiche, of Alfeld, from the northern part of the Transvaal. The first specimen (in 1890) went to the Zoological Garden of Berlin and the second (in 1890) to the Zoological Garden of Vienna. Both were young males, and generally of a reddish colour, with the horns slightly developed. They did not live long after their arrival in the Gardens.

The Pallah is represented in our National Collection by a mounted male from Kilimanjaro shot by Mr. F. J. Jackson and by a mounted head from Lake Elmetaita presented by Captain Lugard, the horns of which are amongst the longest of known specimens. There is likewise a mounted head from the Zomba highlands presented by Sir Harry Johnston and representing the short-horned race which inhabits the mountain-districts south of Lake Nyasa. Besides these there are skulls, skins, and horns from various districts in South and East Africa.

Our illustration of the Pallah (Plate XLVIII.) has been put upon the stone by Mr. Smit from a water-colour drawing by Wolf prepared for the late Sir Victor Brooke and now belonging to Sir Douglas Brooke. The drawing is noted on the back as having been taken from a head belonging to Mr. Selous and a loose skin. It represents an adult male in two positions. The female, as already stated, is absolutely hornless.

The woodcut (fig. 47, p. 23), which gives a front view of a good head of the Pallah, was drawn by Mr. Smit under Sir Victor Brooke’s directions.

August, 1897.

[25]

78. THE ANGOLAN PALLAH.
ÆPYCEROS PETERSI, Bocage.

Similar, so far as is yet known, to Æ. melampus in all respects except that on the face, as is shown in our woodcut (p. 26), there is a prominent brown patch running along the top of the muzzle. This character is said to be perfectly constant, and we therefore admit for the present the validity of the Angolan form as a distinct species.

The Angolan Pallah was first recognized as a distinct species by Prof. J. V. Barboza du Bocage, a distinguished naturalist of Portugal, in a list of Angolan Antelopes published in the Zoological Society’s ‘Proceedings’ for 1878. M. du Bocage based his description upon two specimens forwarded to the Lisbon Museum by the well-known explorer d’Anchieta. Of these the male was stated to have come from Capangombe, the female from Humbe—two places both in the province of Mossamedes north of the Cunene River. M. Bocage distinguished the new species from Æ. melampus principally by its black face, and dedicated it to the late Professor Peters, of Berlin, whose opinion agreed with his that it was distinct. It is probable that the skull from the Cunene River, obtained by Heer Van der Kellen in October 1885, and referred by Dr. Jentink, in his paper on Mammals from Mossamedes, to Æ. melampus, may belong properly to Æ. petersi.

Fig. 48.

Front view of head of Angolan Pallah.

(P. Z. S. 1890, p. 460.)

In 1889 Capt. F. Cookson, during a sporting excursion into Hasholand or[26] Kaokoland, in the neighbourhood of the Cunene River met with some twenty or more specimens of this Antelope, and brought back a single head to England. This head, mounted by Mr. Rowland Ward, was exhibited by Sclater at a meeting of the Zoological Society on June 17th, 1890, as an example of Æpyceros petersi. The notice of Sclater’s exhibition published in the Zoological Society’s ‘Proceedings’ was accompanied by an illustration, which, by the kindness of the Council of the Zoological Society, we are enabled to reproduce (fig. 48). The dimensions of these horns are given by Mr. Rowland[27] Ward, in his ‘Records of Big Game’ (1896), as 18¾ inches in a straight line and 22¾ on front curve, and the distance between the tips as 12¼ inches.

So far as we know, this is all the evidence to be offered as to the existence of this species, concerning which further particulars would be very desirable. There is no example of it in the British Museum.

August, 1897.

Size medium. Nose large, elongate, bent downwards, and inflated; the nostrils opening downwards. Tail short. Mammæ 4. Accessory hoofs present.

Skull with short nasals and premaxillaries, and an exceedingly large and high nasal opening; small supraorbital pits; no lachrymal vacuities; anteorbital fossæ shallow. Lower premolars two, at least in the recent species.

Horns of medium length, cylindrical, rather irregularly lyrate, strongly ringed, pale whitish or amber-coloured. Female hornless.

One species only.

THE BOOK OF ANTELOPES, PL. XLIX.

Wolf del. J. Smit lith.

Hanhart imp.

The Saiga

SAIGA TATARICA.

Published by R. H. Porter.

[31]

79. THE SAIGA.
SAIGA TATARICA (Linn.).
[PLATE XLIX.]

Height at withers about 30 inches. General colour in summer dull yellowish, with a whitish throat and indistinct facial markings; in winter nearly uniform whitish all over, without markings anywhere. Ears very short, thickly haired. Tail short, uniform in colour with the body.

Skull and horns of male as above described (p. 29). The dimensions of an old male skull are:—Basal length 9·5 inches, greatest breadth 5·1, muzzle to orbit 6·3.

[33]

The horns attain a length of about 13 or 14 inches, and are of a peculiar waxy or pale amber-colour.

Female similar, but without horns.

The Saiga, although closely allied to the Gazelles in structure, is, as will be seen from our figure, very different in external appearance, especially as regards the bloated form of the nose in the adult male, which gives it a most ungainly look and renders it easily distinguishable from all its allies of this group.

The Saiga was known to many of the ancient writers, and is described and figured by Gesner, in his ‘History of Quadrupeds,’ as an inhabitant of Scythia and Sarmatia, under the name “Colus,” which is said to have been formed by transposition from the native name “Sulac.” The earliest good account of it, however, is that of the well-known naturalist J. G. Gmelin, who met with it during his travels in Siberia between 1733 and 1743, and described it at full length, in an article on new quadrupeds published at St. Petersburg in 1760, under the name of “Ibex imberbis.” Upon Gmelin’s Ibex imberbis Linnæus, in his ‘Systema Naturæ,’ based his Capra tatarica. Of the two generic names proposed for this Antelope, Saiga by Gray in 1843, and Colus by Wagner in the following year, we naturally prefer the oldest, and adopt as the proper name of this Antelope, which is the sole representative of its genus, Saiga tatarica.

Buffon, in his ‘Histoire Naturelle,’ also employed Saiga as the name of this animal and based his account of it mainly upon Gmelin’s description, stating, however, that there were specimens of its horns in the Royal Cabinet at Paris. Following the prior authorities, he describes the Saiga as a kind of wild goat found at that epoch in Hungary, Poland, Tartary, and Southern Siberia in herds on the plains, very fleet and active, and difficult of capture. We shall see, however, that the range of this animal in Europe has become very much more restricted in recent times.

The best modern account of the Saiga is that given in 1865, in the Bulletin of the Imperial Society of Naturalists of Moscow, by Herr Constantin Glitsch, of Sarepta on the Lower Volga, who was employed for two years by the Imperial Russian Society of Acclimatisation to obtain living examples of this Antelope for the Zoological Garden of Moscow.

[34]

In the days of Pallas, Herr Glitsch tells us, the Saiga had a wide distribution in Europe, extending from the borders of Poland, all across the Dnieper and the great flat southern portion of Russia to the Caucasus and the Caspian. The European herds of this animal were also often reinforced by large accessions from the steppes of Western Asia, which, driven by stress of famine from their native haunts, crossed the Ural and the Volga by the ice in winter. A hundred years later we find a great change in the range of the Saiga, caused by the increase of cultivation and population in the European portion of its range, which has driven this animal back into the East. On the Dnieper, Herr Glitsch tells us, the Saiga has altogether disappeared, in the Ukraine it is no longer to be found, and even on the Don, where it was formerly so plentiful, it is quite a scarce animal. Nowadays, in fact, in Europe the Saiga is confined to the Kalmuk Steppes between the Don and the Volga, and is found only within the triangle lying between these two rivers, of which Tzaritzyn on the Volga forms the northern point.

On the flat and treeless plains which lie within these limits the Saiga still exists in tolerable abundance, though diminishing in numbers yearly as population increases. In the summer months it is distributed over the whole of this area; in winter, beginning from November, it is driven by the snow and cold from its northern resorts towards the south, where it finds shelter in the rich grassy valleys of the Sal and the Manitsch. Here the Saiga passes the winter on ground generally free from snow. Here it breeds in the spring, and as soon as the snow is melted in the more northern plains it begins its migration to the North. At this season the Saigas go northwards in considerable herds, the bucks first, followed by the does, and by the end of May they have all reached the most northern boundaries of their range. But there are many circumstances which interfere with the regularity of this migration, and at Sarepta, near the north end of their area, there are remarkable variations in their numbers. In some summers only a few scattered individuals are to be met with, in other years large herds are to be found in this district throughout the summer. But in very severe winters, when even the most southern districts inhabited by this Antelope are invaded by excessive cold and deep snow, the hungry beasts are driven all over the country in search of food, and stray even as far north as the vicinity of Sarepta. On these occasions whole herds are often entombed in the snow-drifts and fall an easy prey to the natives, who follow them on horseback and slaughter[35] them by hundreds. Under these circumstances it can easily be understood that the Saiga is a gradually vanishing animal in Europe. One thing, however, is in their favour, that the males, whose presence is betrayed by their horns, fall more easy victims to the hunter than the hornless females, which are more readily concealed in the herbage and thus escape notice.

Fig. 49.

Group of Saigas (1/12 nat. size).

(From the ‘Royal Natural History,’ vol. ii. p. 298.)

Herr Glitsch gives us detailed and excellent descriptions of the form and colouring of the Saiga, and of the other peculiarities of the animal of both sexes and in all ages. In the winter coat the hairs on the upper part of the body are from two to three inches long, rather shorter on the underparts, and a long beard extends from the chin down the middle line of the neck to the breast. The older the animal is the brighter is its winter dress.

[36]

The voice of the Saiga is stated by Glitsch to be a deep loud bleat, which is frequently uttered by the young animals, but by the older animals only in the pairing-season and when they are wounded. The hearing, the sight, and the smell of the Saiga are all highly developed, and combine to render it a very difficult animal for the hunter to approach.

The Saigas are said to begin breeding about the middle of December (new style), and at this season commonly assemble in large herds in the warm side-valleys of the Sal and Manitsch, which are mostly free from snow. At this time the young are said to be driven away from the parents in flocks into the thickets, while the males fight fiercely one with another for the possession of the females. The female is stated to go five months with young, and to bring forth about the middle of May amongst the higher vegetation of the steppe. As a rule, she produces two young ones, seldom only one. The mother is sometimes seen followed by three young ones, but in such a case the third is, probably, an adopted animal. In the morning, after suckling her young ones, the mother leaves them concealed in the herbage, and goes far off to feed, returning to them only in the evening and staying with them all night. In about four weeks’ time the young Saigas learn to feed themselves, and the young horns begin to appear in the bucks. They suck, however, till the end of October, and follow after the mother up to the winter. The food of the Saiga consists not so much of the true grasses as of the leafy shrubs of the steppes, such as Artemisia, Atriplex, and Glycirhiza, as well as Inula dysenterica and other saline plants.

Besides mankind, Herr Glitsch tells us, the Saiga Antelope in the Volga district has no special enemy. The wolves and foxes, the only large beasts of prey of these steppes, can only attack quite young animals, the older ones easily making their escape. They have one great plague in the steppes, however, in the insects, especially a species of Œstrus, by which at times they seem to be driven nearly crazy, and with the eggs and larvæ of which their skins seem to be almost always infested.

The flesh of the Saiga is said to be particularly tender and well-flavoured, and more like good mutton than anything else.

The favourite mode of chase of the Saiga is to drive out on to the steppes at early dawn with a cart containing provisions, and, after hiding the cart in some ravine, to stalk them with a rifle in the same manner as other large game-animals. But they are also occasionally taken in steel traps which are[37] set upon their favourite runs. The Kalmuks use leather slings for the same purpose.

Beyond the Ural River the Saiga extends widely over the Kirghiz Steppes of Central Asia north of the Aral. Mr. William Bateson, F.R.S., has kindly favoured us with the following notes of what he heard and saw of the Saiga when in this district in 1896–7:—

As regards the range of the Saiga at the present time, Herr E. Büchner, Director of the Zoological Museum of the Imperial Academy of Sciences of St. Petersburg, has kindly favoured us with the following particulars:—

The Saiga is still met with, although very unfrequently, in the country of the Ural Cossacks between the Wolga and the Ural, and extends occasionally into the Government of Samara. East of the river Ural its range extends over the Kirghiz Steppes and the steppe district of all West Siberia—Turgai, Akmolinsk, and Semipalatinsk. South of this the Saiga is also found in the steppes of Russian Turkestan and in the Dsungarian steppes of Western Mongolia, but not in Transcaspia.

[38]

Such is the range of the Saiga at present. As already shown, it was much wider than now even within the period of history. But when we go back into the Pleistocene times we have good evidence that the Saiga had a very much more extensive range, its fossil remains having been obtained from the caverns and superficial deposits of Hungary, Belgium, and Southern France. In the last-named country the researches of French palæontologists have proved that its bones and teeth occur in considerable numbers in certain of the cave deposits in the Departments of Vienne, Dordogne, Tarn-et-Garonne, and Haute-Garonne. Moreover, as shown by Mons. Gervais, at least one recognizable sketch of the head of the Saiga has been found on an artificially incised bone of the character so often met with in caverns where relics of human handiwork occur. It appears, therefore, that the Saiga inhabited Western Europe as late as the era of Palæolithic man, and was, moreover, in all probability one of the objects of his chase.

Still more interesting, however, is it to find that, as shown by Mr. A. Smith Woodward in a paper read before the Zoological Society in 1890, the Saiga was also found in former days in Great Britain. During excavations made in that year in the Pleistocene deposits near Twickenham, a fine example of the frontlet and horn-cores of an adult male Saiga tatarica was discovered. By the kindness of the Zoological Society we are enabled to reproduce the figure of this interesting specimen (fig. 50, p. 39), which was exhibited by Mr. A. Smith Woodward on the occasion in question, and is now in the gallery of the British Museum.

Finally we may mention that, as has been recorded by Prof. Nehring, there have been discovered in Moravia remains of a Saiga differing from the living species in having three, in place of two, lower premolars[2]. From the occurrence of these remains, and those of other mammals now characteristic of the steppes in Western Europe, it has been argued by geologists that steppe-like conditions and climate must formerly have prevailed over large districts that have now quite changed their character.

Fig. 50.

Frontlet and horns of Saiga (fossil), ♂. ½ nat. size.

(P. Z. S. 1890, p. 614.)

The Saiga has occasionally, but not often, been brought alive to the menageries of Western Europe. In 1864 and 1865 young male specimens of this Antelope were first received from Moscow by the Zoological Society of London. In November 1866 a pair of Saigas was deposited in the[39] Zoological Society’s Gardens, and subsequently purchased, after living for several months in the Regent’s Park Gardens. An excellent coloured figure of these strange animals was made by Mr. J. Wolf in 1867, and published in the Society’s ‘Proceedings,’ and after their death Dr. Murie, then Prosector to the Society, based upon them an elaborate account of their structure and anatomy, which will be found in the volume of the same publication for the year 1870. By the kindness of the Zoological Society we are enabled to reproduce here an excellent figure of the head of the adult male Saiga in its winter coat, taken from a drawing made by Mr. Berjeau under Dr. Murie’s[40] supervision. We cannot do better than refer those who are interested in the structure and anatomy of the Saiga to Dr. Murie’s excellent article, from which, however, we venture to borrow his account of the cutaneous glands of this curious form, which appear to be not less than ten in number.

Fig. 51.

Head of male Saiga in its winter dress.

(P. Z. S. 1870, p. 495.)

The Saiga is represented in the British Museum by a mounted pair from Sarepta on the Volga, and by other skins and skeletons from the same locality. There are also some horns obtained by Dr. O. Finsch on the steppe near Saisan, on the Russo-Chinese frontier, in 1876 (see Finsch, ‘Reise nach West-Sibirien im Jahre 1876,’ p. 193).

Our figure of the Saiga (Plate XLIX.) has been put upon the stone by Mr. Smit from a black-and-white sketch prepared by Mr. Wolf for the late Sir Victor Brooke. The original sketch, which belongs to Sir Douglas Brooke, has been kindly lent to us for examination. We regret to say, however, that we have no particulars as to the individual from which Mr. Wolf’s drawing was taken.

August, 1897.

Size medium. Nose less bent downwards than in Saiga, but more swollen laterally, at least in the male. No suborbital glands. Tail short. Mammæ 2. Large glands in feet and groin.

Skull without distinct pits between the eyes, or lachrymal vacuities, or anteorbital fossæ. Nasal opening ample, but not so large as that of Saiga.

Horns long, erect, compressed, slightly diverging, nearly straight below, evenly curving forwards above; ringed in front. Female hornless.

One species only.

THE BOOK OF ANTELOPES, PL. L.

Wolf del, J. Smit lith.

Hanhart imp.

The Chiru.

PANTHOLOPS HODGSONI.

Published by R. H Porter

[45]

80. THE CHIRU.
PANTHOLOPS HODGSONI (Abel).

Height at withers about 31 or 32 inches. Hair very close, thick, and crisp. Colour pale fawn, with a peculiar fulvous or pinkish suffusion, especially on the flanks. Belly whitish, not sharply separated from the colour of the sides. Face of male black, crown and neck whitish. Sides of muzzle in male markedly swollen. Ears short, but pointed, whitish. Limbs pale greyish white, a black line running down their anterior faces in the male; female without blacker markings. Tail short, coloured like the rump.

Skull dimensions of a male:—Basal length 10·2 inches, greatest breadth 5, muzzle to orbit 6·4.

Horns long, very graceful, nearly straight, only slightly curved backwards below and forwards above, remarkably uniform in length and curvature, generally from 23 to 26 inches in length, the largest recorded being just under 28 inches.

Female similar to male, but without horns.

The Chiru, or Tibetan Antelope as it is often called, although known by the vague reports of the natives as long ago, perhaps, as 1816, was first introduced to science by Abel in 1826, from information and specimens furnished to him by the great naturalist and collector Hodgson, whose name it worthily bears. As we learn from Hodgson’s article published in ‘Gleanings in Science’ for 1830, it was in 1824 or 1825 that a live Chiru was sent to him in Nepal, where he was British Resident at the Court of Catmandu. Hodgson, as was his custom, drew up an elaborate description of the animal, and, after its death, sent the notes along with the skin to[47] Dr. C. Abel, who was at that time one of the Secretaries of the Asiatic Society at Calcutta. Dr. Abel, after making a few additions to the description, and proposing to name the animal Antilope hodgsoni, read his paper at one of the Meetings of the Asiatic Society, and, as it appears from notices in the ‘Philosophical Magazine’ of 1826 and ‘Brewster’s Journal of Science’ of 1827, had it published in the Calcutta Government Gazette or Journal.

But Hodgson, probably owing to the death of Dr. Abel shortly afterwards, was unaware of this fact, and believing that Dr. Abel had lost or neglected his communication, redescribed the species in 1830 under the name Antilope hodgsoni, which he was told that Dr. Abel had applied to it. At that date (1830) Hodgson states that the living specimen already referred to was the only example he had ever seen of this animal, and that up to that time he had never been able to get another example of it alive or dead. It is clear, however, that Hodgson shortly after this date was enabled to obtain further specimens of this Antelope. In one of his letters published in the ‘Proceedings of the Zoological Society’ for 1832 it is stated that three individuals had been examined, and in a subsequent communication (dated from Nepal in February 1834) skins of the Chiru of both sexes are referred to as being amongst other skins of mammals and birds which had been recently despatched to the Society. In the latter communication also Mr. Hodgson suggests the propriety of regarding the Chiru as representing “a new subgenus to be termed Pantholops, the vulgar old name for the Unicorn.” Naturalists have generally acquiesced in Hodgson’s suggestion on this point, and we follow the usual practice in denominating the present species Pantholops hodgsoni.

Other names, however, have been proposed. In 1827 Lesson, in his ‘Manuel de Mammalogie,’ called this Antelope Antilope chiru, quoting as his reference an article in the ‘Quarterly Oriental Magazine’ for 1824 (p. 260), which is, however, merely another version of Abel’s paper.

About the same date also Hamilton Smith, in one of the volumes of Griffith’s edition of Cuvier’s ‘Animal Kingdom,’ proposed, with a note of interrogation, to give the name Antilope kemas to the Chiru, quoting its description from one of the above-mentioned reports of Abel’s original paper.

We are quite satisfied, however, that it is best to employ the specific name hodgsoni for this species as that which was first applied to it.

Since the days when Hodgson was Resident in Nepal many British travellers and sportsmen have penetrated into the snowy ranges of the Himalaya, and[48] have met with the Tibetan Antelope. Sir Joseph Hooker, in the second volume of his ‘Himalayan Journal,’ tells us that he saw Chirus on the Cholamoo lakes near the Donkia Pass in Sikim in October 1849. They were feeding in company with “Gaurs” (Gazella picticaudata) upon the short grass about the lake, which lies at an elevation of some 17,000 feet above the sea-level. Sir Joseph Hooker gives an excellent figure of the remarkable horns of this Antelope, which by his kindness we are enabled to reproduce, and alludes to the ideas of Hodgson (which were shared in by Hue and Gabet) of the profile view of these horns having given rise to the belief of the existence of a Unicorn in Tibet. We should mention that Blanford when he visited Sikim in 1871 was told by the Tibetans that the Chiru is not now found within a long distance of the frontier, but only beyond it in Tibet proper. He admits, however, that it is not probable that there could have been any mistake about so fine and conspicuous an animal.

Fig. 52.

Horns of Chiru.

(From Hooker’s ‘Himalayan Journal,’ vol. ii. p. 158.)

[49]

But by far the most complete account of the Chiru yet published is that given by General Kinloch in his excellent volume on the ‘Large Game of Tibet and Northern India,’ from the second edition of which, published in 1885, we venture to extract the following particulars:—

How far the Chiru extends into the high plateau of Northern Asia beyond the Himalayas it is yet a little uncertain. Dr. Blanford, in his account of the mammals collected by Stoliczka during the Second Yarkand Mission (where excellent coloured figures of both sexes of this Antelope are given), tells us that it has been found in the Kuen-lun range, but has not been met with further north-west or west. It is also, as we are told by the great Russian traveller and naturalist Przewalski, a characteristic animal of the highlands of Northern Tibet. The “Orongo,” as it is here called by the Monguls and Tanguts, was first met with by the great traveller after crossing the Burkhan Buddha range, beyond which it was found distributed to the south as far as the Tang-la mountains. In Mr. Delmar Morgan’s translation of Przewalski’s travels will be found the following passages relating to the habits of this animal, of which, in the original Russian edition of the work, both sexes are figured:—

It is almost unnecessary to say that living specimens of the Chiru have never, as yet, been brought to Europe.

The British Museum contains a mounted specimen of an adult male of the Chiru, obtained by Mr. Mandelli in Sikim and presented by Dr. W. T. Blanford; also some specimens presented by Hodgson, and a number of very fine skulls and horns from Ladakh and Kumaon from the Hume Collection.

[52]

Our illustration (Plate L.), which represents a male of this animal in a snowstorm, has been put upon the stone by Mr. Smit from a coloured drawing prepared by Mr. Wolf under the directions of the late Sir Victor Brooke.

August, 1897.

General characters as in Gazella, but, as in Saiga tatarica alone of Ruminants, with only two lower premolars, and the upper anterior premolar reduced to half the size of the second. Back with a peculiar elongate evertible fold in the skin.

Skull with small but particularly deep anteorbital fossæ, no anteorbital vacuities, and very broad and open posterior nares.

Horns medium, lyrate, twisted inwards, with a double serpentine curvature, convex inwards and in front below, outwards and behind above. The points turned inwards or backwards.

[54]

THE BOOK OF ANTELOPES, PL. LI

Wolf del J Smit lith

Hanhart imp

The Springbuck

ANTIDORCAS EUCHORE.

Published by R H Porter

[55]

81. THE SPRINGBUCK.
ANTIDORCAS EUCHORE (Zimm.).
[PLATE LI.]

Height at withers 31 or 32 inches. General colour bright rufous fawn; a strongly marked, dark, lateral band present, as in many Gazelles. Face pure white, a narrow fawn-coloured line running forwards to the muzzle from the openings of the anteorbital glands on each side. Crown and centre of forehead fawn-coloured like the neck and back. Ears long and pointed, their backs white or pale fawn. Posterior back with a strongly contrasted pure white line, the white hairs of which are placed in a fold of the skin, which fold is everted when the animal is excited, and then forms a prominent white[57] crest; rump white, in continuation with the dorsal line; tail also white basally, black and crested terminally. Belly pure white. Limbs fawn-coloured externally, white on their inner sides and behind.

Skull-dimensions of a male:—Basal length 7·8 inches, greatest breadth 3·7, muzzle to orbit 4·7.

Horns attaining a length of about 14 or 15 inches round the curves, one specimen being recorded as long as 19 inches.

Female similar to male, but horns smaller and not so strongly ringed at the base.

The Springbuck is, no doubt, very closely allied to the Gazelles; but in view of its peculiar dentition, which, as we have pointed out above (p. 53), is unique in the bovine family, and of the remarkable dorsal fold of skin, which is not found in any of its allies, we have thought it advisable to adopt for it the generic term Antidorcas, first provisionally suggested for it by Sundevall in 1847, and subsequently employed by many naturalists. The locality of the Springbuck is also quite distinct from that of the typical Gazelles, which are essentially a northern group, no true Gazellæ being met with until we advance as far north as German East Africa.

This Antelope, with its bright colour and lively movements, as may be easily imagined, quickly attracted the notice of the early Dutch settlers at the Cape and received from them the appropriate name of “Springbok,” from the extraordinary springs and leaps which it makes in running. The first scientific account of it published appears to be that given by Allamand in Schneider’s edition of the ‘Histoire Naturelle’ of Buffon, published at Amsterdam about 1778. In the fourth volume of the ‘Supplement’ of this rather rare work, for the privilege of consulting which we are much indebted to Sir Edmund Loder, will be found (under the head of an addition to the article on Gazelles issued in the twelfth volume of the original work) described and figured “La Gazelle à bourse sur le dos,” as Allamand named the Springbuck. Allamand informs us that his figure and description (which unmistakably relate to this animal) were taken from a specimen then living in the menagerie of the Prince of Orange, which had been brought from the[58] Cape by Capt. Gordon, and was the only survivor of twelve examples of this animal with which Captain Gordon had started for Europe.

Upon Allamand’s “Gazelle à bourse” Zimmermann, in the second volume of his ‘Geographische Geschichte,’ issued in 1780, established his Antilope marsupialis, adding a Latin diagnosis and a shortened translation of Allamand’s description. In the meantime, however, another name seems to have been proposed for the same animal by Forster, who, as we are informed by Zimmermann in the third volume of his ‘Geographische Geschichte,’ had called the Springbuck Antilope euchore. This no doubt was done in the famous ‘Descriptiones Animalium,’ which, although generally accessible in manuscript to the naturalists of the day, and frequently quoted by them, was not published until 1844. However, as Forster’s name for the Springbuck has been accepted nearly universally by subsequent naturalists, we do not now propose to change the name by which this animal has been known for so many years.

The immediately succeeding writers added little or nothing to our knowledge of this Antelope until about 1829, when Lichtenstein, in the second number of his ‘Darstellung der Thiere,’ gave coloured figures of both its sexes under the name “Antilope euchore, Forster,” from specimens in the Berlin Museum procured by him or his assistants in Cafferland.

A few years later Cornwallis Harris visited South Africa. In his great work on the results of his journey subsequently published, this celebrated sportsman and naturalist devotes his third plate to the illustration of a group of Springbucks, which he describes at the period of his writing (1840) as then still abundant in the Colony and “distributed over the arid plains beyond it in unlimited herds.” “Amongst the many striking novelties,” Cornwallis Harris writes, “which present themselves to the eye of the traveller in Southern Africa there are, perhaps, few objects more conspicuous or more beautiful than the dancing herds of graceful Springbucks which speckle the broad plains of the interior.”

Although not still met with in the countless thousands described by Cornwallis Harris, the Springbuck, we are pleased to be able to say, is even now abundant in many parts of the Cape Colony, and Springbuck shooting is still one of the recognized sports of its inhabitants and of visitors to Southern Africa who go in search of game. Mr. H. A. Bryden, in his well-known volume ‘Kloof and Karroo,’ devotes a whole chapter to the delights of Springbuck shooting, and tells us that of late years large tracts of waste land in the Colony have been fenced in in order to preserve these Antelopes. For example, as the ‘Graaf Reinet Advertiser,’ of November 1886, informs us, Shirlands, the property of Mr. John Priest, of that district, was, twelve to thirteen years ago, a piece of waste land abandoned to squatters. Now there are 16,000 morgen (more than 32,000 acres) fenced in with wire. Within this fence there are fully a thousand Springboks where formerly only a few remained “harassed and hunted to death by impoverished lazy squatters.”

In the Cape Colony Mr. W. L. Sclater, the Director of the South African Museum, Cape Town, kindly informs us that in the west of the Colony the Springbuck is met with in Namaqualand, Clanwilliam, Beaufort West, Prince Albert, and the adjoining districts. In the middle of the Colony it is found in Uitenhaag, Graaff Reinet, Colesberg, Albert, and Queenstown, but is rare in East Albany. On the north it occurs in Great Namaqualand and Damaraland, also in Kimberley, Barkly West, and Herbert. But it must be understood that it is mostly confined in all these districts to those farms of the Dutch and English settlers where it is preserved, and that permission to shoot it must on all occasions be obtained. The same is the case in the Orange Free State and Transvaal. In Bechuanaland, being wholly unprotected, the Springbuck has in recent years been much shot down, except on the open arid flats north and south of the Botletle and the neighbourhood of the great Makari-kari Salt-pan, where Messrs. Nicolls and Eglington say it still roams in large herds.

[61]

As regards the northern limit of the Springbuck, it certainly does not cross the Zambesi in any place so far as we have been able to ascertain.

Fig. 53.

Horns of Springbuck, ♂ and ♀.

Mr. F. C. Selous (P. Z. S. 1881, p. 757) says that its northern range is bounded on the east by the thick forests which run east and west south of the Mababe River. Westwards, as already stated, it occurs in the district of Lake Ngami and throughout Damaraland up to the Portuguese province of Mossamedes, whence specimens have been forwarded to the Lisbon Museum by their energetic collector M. d’Anchieta.

[62]

Writing quite recently to Sclater, Mr. J. ffolliott Darling gives the following notes:—

White and Albino varieties are not so frequently met with amongst the Bovidæ as in some other groups of mammals. But the ‘Johannesburg Times’ of January 22nd, 1897, informs us that a perfectly white Springbuck, caught in the Orange Free State, and supposed to be about eleven months old, was at that time being exhibited in Johannesburg by Messrs. Colquhoun and Hill, of Jeppe Street. Such a novelty as a white Springbuck was previously quite unknown in the Transvaal. This communication was sent to us accompanied by a photograph of the animal taken from life, from which it would appear that its colour was absolutely of a spotless white.

The Springbuck, although not unfrequently seen in the Zoological Gardens of Europe, is, as might be supposed, from its free and active habits, somewhat[63] impatient of captivity and does not thrive in confinement except in occasional instances. The Zoological Society acquired their first specimen (by purchase) on July 9th, 1852, and, as will be seen by reference to their published Lists of Animals, others have been subsequently received at short intervals since that date. At the time we are writing there is a fine pair in the Society’s Gardens, deposited by H.R.H. the Prince of Wales in March, 1893, which are still doing well. Dr. Wünderlich, Director of the Zoological Garden of Cologne, has kindly furnished us with notes upon a pair of this Antelope which he bought on the 13th May, 1896, from Herr Reiche, of Alfeld. They bred on the 24th May last year, and after a period of 171 days a young one of the female sex was born on the 12th November last. The young one at birth was 45 cm. (about 17¾ English inches) in height, and generally of a yellowish-grey colouring. The side stripe was rather darker, but by no means so clearly defined as it is in the adult animal. The under surface and inner sides of the limbs were white, as in the adult. On the face a dark stripe from the eye to the corner of the mouth was visible, but the cheeks and chin generally were yellowish grey like the sides of the body. The little animal did well in company with its mother at first, and after 15 days began to eat corn. Unfortunately, however, it did not continue to thrive, and died on December 21st, when about 40 days old.

The flesh of the Springbuck is much esteemed by the epicures of the Cape Colony, and has been occasionally brought to London in a refrigerator for consumption here. In the ‘Field’ for 1892 (vol. lxxx. p. 390) will be found an account of its successful importation by Messrs. Brooks, of Leadenhall Market, and of the high appreciation it met with by those who tried the “venison,” which was pronounced to be “in good condition, not the least high, and tasting not unlike Chamois.”

Our figure of the adult male Springbuck, with a herd of these animals in the distance, has been prepared by Mr. Smit from an original sketch by Wolf, which is now in the possession of Sir Douglas Brooke, and has been kindly lent to us for examination.

There is a good stuffed specimen of an adult male Springbuck in the Gallery of the British Museum. It was obtained by Mr. F. C. Selous at Mahemfontein, in the Orange Free State, in 1896, and was presented to the collection by that gentleman. From a pencil-note on the back of Mr. Wolf’s sketch it would appear to have been taken from a specimen of the head of[64] this species formerly exhibited in the same Museum. The National Collection has also some skins, skulls, and horns of the Springbuck from various parts of South Africa; but good skulls of this Antelope are still desiderata to the National Collection, and a series of them, with dates and localities, would be much appreciated.

August, 1897.

Size medium, but with a considerable range of variation. General form normal; the muzzle simple, neither expanded as in Pantholops nor elongated as in Saiga; the neck of ordinary length, and the back without any evertible fold as in Antidorcas. Coloration ordinarily sandy, with a white belly, the face generally marked with dark and light streaks; streaks also generally present on the flanks and rump. Knee-brushes usually present. Tail short or of medium length.

Skull generally with shallow anteorbital fossæ corresponding to the anteorbital glands, but occasionally—in the procaprine section—without any trace of them. Premolars 3/3, as usual in the Bovines.

Horns generally present in both sexes, the females of the first four species alone being without them. In the male the horns are strong, prominently[66] ribbed, and generally of medium length, about the length of the head, but occasionally considerably longer. With the exception of their tips they are curved backwards, so as to be convex forwards below, while their ends are commonly more or less recurved forwards or upwards. The degrees of curvature seem to be fairly constant within the species, and to afford very fair specific characters. On the other hand, in the female the horns are slender, straighter, and shorter than in the male, very variable in direction, and as a rule showing little of the characteristic curvature peculiar to the male of each species, although there is a certain correspondence between the lengths of the horns in the two sexes.

The genus Gazella contains the great majority of the members of the present subfamily, and forms a very natural and easily defined group. All the species are lightly built and delicate animals, and are among the best known of all the Antelopes, on account of their beauty and the fact that they are common in confinement, so that every zoological garden is always well provided with examples of them. In the Zoological Society’s Gardens at the present time no less than ten species are represented.

The genus Gazella was always a favourite one with Sir Victor Brooke, who devoted much time to its elucidation, and published in 1873 a monograph of it, which up to the present time has been the standard work on the subject. From this monograph we venture to quote the following paragraphs which explain the nomenclature of the characteristic markings of the Gazelles: they also give an indication of the difficulties to be met with in working out a genus which, while the largest contained in the present work, is remarkable for the close resemblance of the different species to one another and for the absence of characters which will enable them to be readily separated:—

Since Sir Victor Brooke wrote his monograph of the Gazelles, many species only known to him by descriptions or by imperfect specimens have become represented in our National Museum by complete examples, while several additional species have been discovered. Our arrangement of the species is therefore necessarily different from his, but is, we fear, still very far from being perfect, as more and better specimens of most of the forms are still wanted before their exact geographical distribution, their extent of variation, and their true relationships to each other can be satisfactorily worked out.

As already noted, we have removed from the genus Gazella, under the name Antidorcas, the South-African Springbuck, which Sir Victor Brooke included in it. This being eliminated, the 25 species which we are prepared to recognize as distinct may be arranged as follows:—

• A. Tail quite short. No Gazelline face-markings. Females without horns.
• a. Horns strongly curved backwards. Skull 7 in. or less in basal length.
• a¹. Horns not hooked at tip. (Tibet.)  82. G. picticaudata.
• b¹. Horns hooked at tip. (Mongolia.)  83. G. przewalskii.
• [68]
• b. Horns but little curved backwards, not hooked at tip. Skull about 9 in. in basal length. (N. China.)  84. G. gutturosa.
• B. Tail of average length, its terminal half generally more or less crested with black. Face-markings present. Females (except in G. subgutturosa) with horns.
• a. Upper part of face white (at least in old age), interrupting the central facial band.
• a¹. No horns in female: size larger. (Central and Western Asia.)  85. G. subgutturosa.
• b¹. Horns present in female: size smaller. (Arabia.)  86. G. marica.
• b. Central dark facial band uninterrupted by white above. Horns present in the female.
• a¹. Dark colour of back not invaded by white of rump.
• a². Dark lateral band indistinctly marked, not strong and blackish.
• a³. Tip of horns slightly curved inwards or upwards, not bent in to a right angle.
• a⁴. Horns of medium length.
• a⁵. Horns truly lyrate, the middle portion twisted outwards, the tips reapproaching each other. (Algeria, Egypt, Palestine.)  87. G. dorcas.
• b⁵. Horns not truly lyrate, more or less evenly diverging upwards.
• a⁶. Top of muzzle ordinarily with a black spot on it.
• a⁷. Nose simple.
• a⁸. Larger: hair rough. (Algeria.)  88. G. cuvieri.
• b⁸. Smaller: hair smooth: darker. (Arabia.)  89. G. arabica.
• c⁸. Smaller: hair smooth: lighter. (India and S. Persia.)  90. G. bennetti.
• b⁷. Nose with a flabby corrugated elevation on it. (Somaliland.)  91. G. spekei.
• b⁶. Top of muzzle without a black spot. (Somaliland.)  92. G. pelzelni.
• b⁴. Horns long and very slender. Colours very pale. (Algeria and Egypt.)  93. G. leptoceros.[69]
• b³. Tip of horns hooked inwards or upwards, nearly or quite to a right angle.
• a⁴. General colour pale fawn, the lateral band and other markings also fawn. (Nubia.)  94. G. Isabella.
• b⁴. General colour brownish fawn, the lateral band blackish. (Muscat.)  95. G. muscatensis.
• b². Dark lateral band black and strongly marked.
• a³. Horns abruptly hooked inwards at the ends. (Abyssinia.)  96. G. tilonura.
• b³. Horns not abruptly hooked at end.
• a⁴. No nose-spot; face uniform rufous.
• a⁵. Size smaller. (Senegambia.)  97. G. rufifrons.
• b⁵. Size larger. (Algeria.)  98. G. rufina.
• b⁴. Nose-spot black. (Masai-land.)  99. G. thomsoni.
• b¹. Dark colour of back more or less invaded by white of rump.
• a². A dark pygal band present.
• a³. Dark colour of back continued in middle line on to the top of the tail. Size smaller. (E. African Coast.)  100. G. petersi.
• b³. Dark colour of back shut off from tail, which is enclosed in the white anal disk. Size larger.
• a⁴. Dark lateral bands obsolete, or, in youth, present below the light lateral band only. (Interior of East Africa.)  101. G. granti.
• b⁴. Dark bands present both above and below the light lateral band, uniting with each other behind. (Northern British E. Africa.)  102. G. notata.
• b². No dark pygal band.
• a³. Tail black-tipped. Horns hooked inwards. (Abyssinian Coast and Somaliland.)  103. G. soemmerringi.
• b³. Tail all white, or merely tipped with fawn. Horns hooked upwards and forwards.
• a⁴. Neck and anterior back alone rufous; lines of demarcation indistinct. (Kordofan.)  104. G. ruficollis.[70]
• b⁴. Rufous extending over body and flanks, well defined from the white.
• a⁵. Sides of thighs white, the rufous of body not joining that of hind legs. (Senegal.)  105. G. dama.
• b⁵. Sides of thighs and legs rufous, continuous with that of body. (Morocco.)  106. G. mhorr.

THE BOOK OF ANTELOPES, PL. LII

Wolf del Smit lith

Hanhart imp

The Tibetan Gazelle.

GAZELLA PICTICAUDATA.

Published by R. H. Porter

[71]

82. THE TIBETAN GAZELLE.
GAZELLA PICTICAUDATA (Hodgs.).
[PLATE LII.]

Height at withers about 25 inches. Fur close and thick. General body-colour pale fawn, darkening posteriorly and becoming almost rufous brown along the edges of the white anal patch. No lateral nor pygal bands. Face without any trace of the ordinary Gazelline markings, coloured like the body, or the top of the muzzle sometimes brown; hairs on the sides of the muzzle[72] elongated, so as to form a sort of lateral tuft, which extends backwards under the eyes. Ears short, narrow, pointed, well haired, coloured like the body. Rump with a prominent white patch surrounding the base of the tail. Tail quite short, projecting little beyond the fur, its end black or dark fawn. Limbs white or very pale fawn; no knee-tufts.

“In the summer the coat is short and of a slaty grey colour” (Brooke).

Skull rather broad in proportion to its length. Anteorbital fossæ practically obsolete. Nasals broad behind, evenly tapering forwards. Basal length 6·4 inches, greatest breadth 3·75, muzzle to orbit 4·0.

Horns slender, of median length, much compressed laterally, very closely ringed. With the exception of their tips (2–3 inches), after starting vertically, they curve evenly and strongly backwards, diverging laterally but little. Tips gently curved upwards and slightly inwards, reapproaching each other above to a certain extent.

Female. Similar to the male but without horns.

Like many other Himalayan and Tibetan animals this Antelope first became known to science from the researches of the great Indian naturalist and antiquarian, Bryan Houghton Hodgson, British Resident at the capital of Nepal. Hodgson described it in 1846, in the ‘Journal of the Asiatic Society of Bengal,’ as Procapra picticaudata, and gave a very recognizable figure to accompany his letterpress. He wrote of it as follows:—

Hodgson also entered into the structural peculiarities of this Antelope, which he described at full length. It is quite evident that, as pointed out by him, the present Gazelle, as also the two allied species (G. przewalskii and[73] G. gutturosa), present certain points of difference from the rest of the group, and that there was, therefore, some justification for Hodgson’s proposal of the generic term “Procapra,” although we do not think it necessary to use it. These three species agree among themselves in the females not possessing horns, in the absence of anteorbital glands, and the corresponding absence of a fossa in the skull, in having no brushes on the knees, and in several other characters, which show that they are really more closely connected to each other than to the more typical Gazelles. Nevertheless we think that, on the whole, it is best to include them in the genus Gazella, as no one of these characters is absolutely confined to them. Thus G. subgutturosa, often, though wrongly, placed with them, has no horns in the female, while in other characters it is a true Gazella, and several species besides these three are without knee-brushes, while the anteorbital fossæ in others are so shallow as to be practically non-existent.

Fig. 54.

Skull and horns of the Tibetan Gazelle.

(P. Z. S. 1867, p. 245.)

Soon after his discovery of the Goa, Hodgson forwarded specimens of it to the British Museum, and the species was included in Gray’s catalogues as Procapra picticaudata. Under this name also Gray figured a skull and pair of horns of this Gazelle in 1867, in order to point out its differences from the[74] allied Asiatic form, Gazella gutturosa. This figure (fig. 54, p. 73), by the kind permission of the Zoological Society of London, we are now able to reproduce.

In October 1849, Sir Joseph Hooker, as related in his ‘Himalayan Journals’ (ii. p. 157), met with the Goa feeding on the short grass near the Cholamoo Lake in Sikim, at an elevation of 17,000 feet above the sea-level, and in other adjoining localities on the Donkia Pass between Sikim and Tibet. Through his kindness and that of his publishers we are enabled to introduce the illustration of this striking scene (fig. 55) prepared for his well-known work.

Fig. 55.

Goa Antelopes on the Donkia Pass.

(Hooker’s ‘Himalayan Journals,’ ii. p. 139.)

Other travellers and sportsmen have also noticed the Goa or Tibetan Gazelle in Ladak and on the frontiers of Tibet. But by far the most complete account of the habits and ways of life of this Antelope is that given by Major-General Kinloch in the various editions of his excellent work on Large-Game Shooting in Tibet.

To the east of Ladak, General Kinloch tells us, in the country that lies between the Upper Indus and the Sutlej, are vast expanses of undulating[75] hills and valleys of great elevation utterly destitute of forest and with but scanty indications of vegetation. The greater part of these wild uplands would appear at first to be a perfect desert, but, as a matter of fact, on closer inspection, it will be found that there is hardly a slope, however rocky, or an expanse of sand, however thirsty-looking, where an occasional tuft of grass or bunch of sweet-scented herb may not be found, while wherever streams of water exist their banks are often ornamented by the greenest of turf and studded with flowers of the most brilliant hues. This bleak country, General Kinloch continues, the elevation of which varies from 13,000 to 18,000 feet, is the home of the Goa, which is to be found there scattered about in small parties usually varying from two or three to about a dozen in number, and in certain localities is decidedly plentiful. They are not generally very shy, but will seldom allow the hunter to approach openly within shot.

In 1866 General Kinloch made an expedition to the Tsomoriri Lake in this district, mainly with a view of hunting the Goa. We subjoin an account of his adventures, extracted from his work:—

Besides the experiences of the travellers and sportsmen from the Indian side, the only published record concerning the Tibetan Gazelle, so far as we know, is that of the great Russian explorer Przewalski, who, after treating of the “Orongo” of Northern Tibet (Pantholops hodgsoni), mentions the present species as being found in the same district, and there known to the Mongols as “Ata-dzeren,” or Little Antelope[4]. Przewalski, who met with this animal near the headwaters of the Tatong-gol, in Northern Tibet, and, as he believes, also in the highlands of Kan-su in China, describes its habits as follows:—

There are many specimens of this Antelope in the British Museum presented by Mr. Hodgson, amongst which is the type of the species. There are also in the National Collection skulls from Kumaon and other localities presented by Mr. A. O. Hume, C.B.; from the Changchenmo Valley, Ladak, presented by Mr. R. Lydekker; and from the confines of Tibet north of Sikim, collected by Mandelli and presented by Dr. W. T. Blanford, F.R.S.

There is also a good series of specimens of this Gazelle in the Museum of the Academy of Sciences of St. Petersburg, obtained by the Russian explorers in Northern Tibet and in the mountains of Nan-shan.

Our figure (Plate LII.), which has been put on the stone by Mr. Smit from a sketch prepared for Sir Victor Brooke by Mr. Wolf, represents a male and two females of this species, probably the specimens in the British Museum.

January, 1898.

[78]

THE BOOK OF ANTELOPES PL. LIII.

J Smit del, et lith.

Hanhart imp.

Przewalski’s Gazelle.

GAZELLA PRZEWALSKII.

Published by R H Porter

[79]

83. PRZEWALSKI’S GAZELLE.
GAZELLA PRZEWALSKII, Büchn.
[PLATE LIII.]

Size rather greater than in G. picticaudata. General colour deep fawn in summer, pale finely grizzled fawn in winter. Sides of neck (at least in winter) and top of muzzle slaty brown; no ordinary gazelline face-markings. Ears short, acutely pointed[5]; coloured like the back. Rump with the white of the anal region running up on to the upper surface, divided in its centre by a narrow fawn-coloured line running from the back on to the tail. Tail very short, hidden in the fur; fawn along its top, inconspicuously pale brown at its tip. Front of limbs more or less brownish; no knee-tufts.

Skull short and stoutly built. No anteorbital fossæ. Nasals broad and short; premaxillæ not reaching up to the latter, the nasal opening unusually large and broad. Basal length of an adult male 7 inches, greatest breadth 3·8, muzzle to orbit 4.

Horns of median length and thickness, much compressed laterally; with the exception of their terminal two inches, they are evenly curved backwards[80] and divergent outwards, the divergence increasing above; tips abruptly hooked inwards and slightly upwards, at a sharper angle with the rest of the horn than a right angle.

Female. Similar to the male, but without horns.

Przewalski’s Gazelle, which has been most appropriately named after the famous explorer who discovered it, was first described and figured by Przewalski himself in 1876, but erroneously confounded with the allied form G. gutturosa. Twelve years later, it appears, Przewalski discovered his error, and proposed to rename the animal Antilope cuvieri. But this specific term properly belongs to another species which had been described by Ogilby many years previously. Under these circumstances Dr. Büchner in his account of the Mammals of the Kan-su Expedition of Messrs. Potanin and Beresowski, proposed that this Gazelle should in future be known as Gazella przewalskii—Przewalski’s Gazelle.

In the English translation of Przewalski’s ‘Mongolia’ the habits of this species are described as follows:—

Besides the Russian explorers already mentioned, the only traveller, so far as we are aware, that has met with Przewalski’s Gazelle in its native wilds is the well-known explorer Mr. St. George Littledale, F.R.G.S., who brought home a skin and skull of this species from his adventurous journey across Central Asia in 1893, and presented them to the British Museum. In the narrative of Mr. Littledale’s expedition, which is contained in the third volume of the ‘Geographical Journal’ (p. 465), will be found an allusion to this Antelope as observed by him near the Lake Koko Nor. The north shore of this lake, first seen by Mr. Littledale on the 3rd of August, 1893, was flat and swampy, and there were many of these antelopes feeding on it in company with Wild Asses (Equus kiang). In some MS. notes with which Mr. Littledale has kindly favoured us on this subject it is stated that he first saw examples of this Gazelle south of the Nan-Shan mountains in about lat. 38° 30´ N. and long. 96° 30´ E. On that occasion, he says, they were high up above the party, and nearly all males, but, as Mr. Littledale was then expecting an attack from the Tanguts, he did not like to leave the caravan to try after them. As the valley of the Buhain-Gol (the river which flows into Lake Koko Nor) was descended, the old males became scarcer, and round the lake, where he procured the specimen now in the British Museum, there were large bands of females accompanied by young males.

Besides Mr. Littledale’s specimens already mentioned, the British Museum[82] contains a beautiful pair of this Antelope obtained in exchange from the Museum at St. Petersburg, from which our figures representing both sexes (Plate LIII.) have been prepared by Mr. Smit. It will be observed at once that though in its general form and coloration this species is somewhat similar to G. picticaudata, the shape of its horns is quite different, and readily distinguishes this species from its allies.

When at St. Petersburg in August 1897, Sclater had the opportunity of examining, under the kind guidance of Herr Büchner, the fine series of specimens of this Antelope in the Zoological Museum of the Imperial Academy of Sciences of that city. The specimens had been obtained in the region of the Koko Nor, in the most northern part of the Chinese province of Kan-su, and in Ordos, which is the country encompassed by the great northern bend of the River Hoang-Ho. In the southern part of this district Przewalski’s Gazelle was met with in great numbers during the Kan-su expedition already mentioned, and many specimens of it were obtained for the Imperial Museum at St. Petersburg.

January, 1898.

THE BOOK OF ANTELOPES, PL. LIV

Wolf del J Smit lith.

Hanhart imp.

The Mongolian Gazelle

GAZELLA GUTTUROSA

Published by R H Porter

[83]

84. THE MONGOLIAN GAZELLE.
GAZELLA GUTTUROSA (Pall.).
[PLATE LIV.]

Size fairly large; height at withers rather over 30 inches; form stout and thick. General colour fawn, paler than in most of the other species, but the rump and sides are white, as is the whole of the limbs. Gazelline face-markings absent; top of the muzzle browner than the rest of the head; sides of the muzzle and cheeks white. Ears short but pointed, thickly furred, their backs pale fawn, nearly white. No lateral bands, either dull or light. No knee-brushes. Tail very short, white, its tip brown.

According to Radde all the parts which are fawn in winter are in summer of an intense isabel-yellow.

Skull long and narrow, with a heavy muzzle. Anteorbital fossæ obsolete. Nasals long and broad. Basal length (in a not fully adult specimen) 9 inches, greatest breadth 4·1, muzzle to orbit 5·5.

Horns short in proportion to the size of the animal; heavily and closely ringed; basally they are parallel to each other, diverging above, with their tips again gently curved in towards each other.

Female. Similar, but without horns.

The great traveller and naturalist, Peter Simon Pallas, whose name we have already often mentioned in the course of this work, was the first technical describer of this Antelope, although he was by no means its discoverer, for he himself quotes previous references to it in the works of older authors. But Pallas, in the Supplement to his memoir on the Antelopes, published 1777, gave us the first scientific description of it, and selected for it the appropriate scientific name gutturosa, by which it has been ever since known. According to Pallas, the first Europeans to become acquainted with this Gazelle were the Jesuit missionaries in China, one of whom, Pereira, as quoted by Witsenius, mentions it as a Chinese animal; while Du Halde, in his great work upon China, describes it, under the name “Hoang-yang” or Capra flava, as wandering about in large flocks in the deserts of Mongolia. Further accounts of this Antelope were subsequently given by Messerschmidt and Gmelin in the Commentaries of the St. Petersburg Academy. These are also quoted by Pallas, who himself met with this animal on the upper course of the River Onon, on the southern frontiers of Transbaikalia. Pallas concludes his history of this species with a lengthened description of its external form and anatomy, and gives an uncoloured figure, in which the peculiar swollen condition of the throat in the male in the breeding-season (whence it was termed gutturosa) is correctly shown.

Pallas’s posthumous work, ‘Zoographia Rosso-Asiatica,’ contains little more than a summary of his previous account of this animal.

The numerous authors who followed Pallas added little or nothing to our knowledge of the Mongolian Gazelle, and were content to base their notices of it almost entirely upon his publications. It is not, in fact, until we come to nearly modern days that we obtain any further original information concerning this animal.

Dr. Gustav Radde, now Director of the Museum at Tiflis, made extensive journeys in South-eastern Siberia, under the patronage of the Imperial Geographical Society of Russia, in 1855 and the three following years, and amassed large zoological collections. One of the volumes of his ‘Reisen im Süden von Ost-Sibirien,’ published at St. Petersburg in 1862, is devoted to an account of the Mammals of South-eastern Siberia, and is, and will long remain, our standard work on this subject. Dr. Radde brought home five good specimens of this Antelope, and commences his account of it with accurate descriptions of its summer and winter pelages. He adds a detailed[86] description of its skull and dentition, and compares them at length with those of Gazella subgutturosa.

As regards its distribution in the present epoch, Dr. Radde points out that, like the Dziggetai (Equus hemionus) and the Argali Sheep (Ovis ammon), the Mongolian Gazelle has retreated to the south and east from the Russian frontiers since the days of Pallas. There are at present only two places on the southern borderlands of Transbaikalia in which this Antelope remains during the summer and breeds every year. One of these is a district east of the Dsŭn-tarei which is seldom entered even by the shepherds of the Cossacks. It is an uninhabited and rather mountainous country, without wood or bushes, varied by salt-and some freshwater lakes, and covered only with yellow Elymus-grasses. The other district, which is of a similar character, lies north of the left bank of the Argunj, where this river enters into the Russian territories between the border-posts of Soktui and Abagaitui.

Dr. Radde gives the following account of the habits of this animal as observed by himself and as obtained from the reports of the natives in 1856:—

Fig. 56.

Skull and horns of the Mongolian Gazelle.

(P. Z. S. 1867, p. 245.)

In 1867 Dr. Lockhart brought home with him from Pekin two skulls of this Antelope and presented them to the British Museum. Dr. Gray read some notes on them at a meeting of the Zoological Society of London in[88] February of that year. These notes were subsequently published in the ‘Proceedings,’ accompanied by an outline figure of one of the heads, which, by the kind favour of the Society, we are enabled to reproduce. Dr. Lockhart gave to Dr. Gray the following information as to this Antelope:—

Besides an adult stuffed specimen of this species in the British Museum, stated, but probably erroneously, to have come from the Kirghiz Steppes, there are several skins and skulls in the collection obtained by the late Consul R. Swinhoe at Pekin, besides the two heads and horns already spoken of as brought home by Dr. Lockhart.

Our illustration (Plate LIV.), which represents both sexes of this Antelope, was put upon the stone by Mr. Smit from a sketch drawn by Mr. Wolf for the late Sir Victor Brooke. We have no record of what were the exact specimens figured.

January, 1898.

THE BOOK OF ANTELOPES, PL. LV.

J. Smit del. et lith.

Hanhart imp.

The Persian Gazelle.

GAZELLA SUBGUTTUROSA.

Published by R. H. Porter.

[89]

85. THE PERSIAN GAZELLE.
GAZELLA SUBGUTTUROSA (Güld.).
[PLATE LV.]

Size medium, height at withers about 26–27 inches. General colour dark sandy fawn. Face-markings indistinct, the central band visible in youth gradually interrupted and replaced by white as age advances. Dark facial streaks in front of eyes present, but little defined. An anteorbital gland present. Larynx swollen, forming a peculiar projection in front of the neck. Ears of medium length, pointed, their backs short-haired even in winter, pale fawn. Dark lateral band not, or scarcely, darker than the back, from which it is separated by an indistinct light lateral band. Pygal band present, not strongly marked. Tail 8 or 10 inches long, crested, black. Knee-brushes present, brown or black.

Skull stoutly built; nasals broad and short; anteorbital fossa well marked. Basal length 8 inches, greatest breadth 3·8, muzzle to orbit 4·6.

[91]

Horns of medium length, thick, evenly diverging from each other as they curve backwards; their tips decidedly, though not abruptly, bent inwards and slightly upwards.

Female similar to the male, but without horns, or occasionally with minute rudiments of them.

The Persian Gazelle, as it is commonly called, is by no means restricted to Persia, but, as we shall presently show, has a wide range through the steppes of Central Asia from the borders of Asia Minor to Northern China. It was first made known to science by Anton Güldenstädt, an enterprising Russian traveller and naturalist of the last century, who met with it in 1772 in the course of his explorations of the countries adjacent to the Black and Caspian Seas. Güldenstädt wrote an elaborate description of it in 1878 in a memoir published two years later in the ‘Acta’ of the Imperial Academy of Sciences of St. Petersburg, and named it “subgutturosa,” “because its throat protruded slightly, but not so much as in Antilope gutturosa.” Pallas, who also observed this Antelope during his travels in Central Asia, included it under Güldenstädt’s name in his ‘Zoographia Rosso-Asiatica.’

After Güldenstädt and Pallas several other Russian naturalists—Hohenacker, Nordmann, and Eichwald—recorded this Gazelle as being met with on the plains of Transcaucasia. Ménétriés, in his memoir on the Zoology of the Caucasus published in 1832, tells us that at that period it was very common, especially in winter, on the vast steppes bordering the Caspian between Baku and Kur, whence, as Herr Büchner has kindly informed us, it extends up the valley of the Kur nearly to Tiflis. Satunin, our most recent authority on the Mammals of this district, states that he found it throughout the steppes of Eastern Transcaucasia, and especially numerous on the Mugan Steppe. Whether this is the Gazelle found on the upper plains of the Tigris and Euphrates, as reported by Danford from hearsay, seems to be uncertain, though it probably penetrates into the highlands of Asiatic Turkey adjacent to Mount Ararat, and is certainly found in the valley of the Araxes.

In Persia, Dr. Blanford tells us, in his volume on the zoology of that country, G. subgutturosa is the Gazelle of the highlands, and is found in[92] almost all the valleys and plains from about 3000 to about 7000 feet above the sea-level, ranging higher in winter and lower in summer, but keeping generally within the limits mentioned. It is unknown in the plains of Mesopotamia, and on the lower ground along the Persian Gulf and the Arabian Sea.

Dr. Blanford adds the following notes from the pen of the late Sir Oliver St. John, who was very well acquainted with Persia and its animals:—

Dr. Blanford has included this Gazelle in the ‘Fauna of British India,’ because, as ascertained by the late Sir Oliver St. John, it occurs in Pishin north of Quetta, now in British territory, though it is not met with in any other part of the Indian Empire.

Throughout the sandy plains along the northern boundary of Afghanistan this Gazelle is abundant. Dr. J. E. T. Aitchison, who accompanied the Commission for the delimitation of the Afghan boundary in 1884, tells us that it was occasionally seen along the whole route from Quetta to Khusan. In the low hills and great gravel plains of the valley of the Hari-rud they were observed everywhere, but were very wary and difficult of approach. In June 1885, at Chinkilok, to the north-west of Herat, between that city and the range of the Parapomisus, Dr. Aitchison caught a young female Gazelle of this species about a day old, and subsequently, on his way home through Persia, obtained three others of about the same age. These four Gazelles, as we have been told, were carried many hundred miles through Persia in large covered baskets on each side of two camels, and were commonly believed by the natives to be Dr. Aitchison’s four wives, the baskets being of the same fashion as those generally used in that country for the conveyance of women! Dr. Aitchison brought his four pets safely home to the Zoological Society’s Gardens, where they throve well and bred in 1887, 1888, and in several succeeding years. Two of the males of this[93] family, born in the Society’s Menagerie in 1892 and 1894, are still living there.

According to Herr Büchner, who has kindly supplied us with some valuable notes on the Asiatic Gazelles, this species is found in suitable localities all over the Transcaspian Provinces of Russia, and ranges northwards to the steppes between the Caspian and the Aral, and eastwards to Lake Balkash. Far beyond this it extends across the southern portions of the great Desert of Gobi into the Chinese Provinces of Zaidam, Alaschan, and Ordos.

On crossing the high range north of the Hindu Koosh into Eastern Turkestan a Gazelle very similar to the Persian Gazelle is met with. Six examples of this form were obtained by the naturalists of the Second Yarkand Mission in 1873–74, and were described by Dr. Blanford in his memoir on the Mammals of that Mission. Dr. Blanford says that it is perhaps a question whether the Eastern Turkestan Gazelle should not be raised to the rank of a species. It differs principally from the typical form in the very much darker markings on the face and in the much smaller degree to which the horns diverge. The size appears rather larger than that of the typical Persian form. But as there are some variations in the face-markings of Persian specimens, Dr. Blanford has considered it better to regard the Yarkand race as only a variety, which he has proposed to call Gazella subgutturosa yarkandensis. Of this subspecies an excellent coloured figure, drawn by Smit, is given in the above-named work. It represents both sexes, and shows the black markings on the face very distinctly.

As pointed out by Dr. Blanford, it is nearly certain that the Gazelle to which Shaw refers, in his volume on ‘High Tartary, Yarkand, and Kashgar’ (page 221), as having been brought to him at Yarkand, and of which he says the Yarkand name is “saikeek,” was of the present species—that is, of the local form Gazella subgutturosa yarkandensis.

We have already mentioned the four living examples of this Gazelle brought to London by Dr. Aitchison and presented to the Zoological Society’s Collection. These, however, were not the first specimens of this animal brought to England alive. As long ago as 1852 females of the present species were obtained from Bussorah on the Persian Gulf and presented to the Society by Alderman Finnis, and in 1869 examples from the same country were given to the collection by the late Mr. T. K. Lynch, F.Z.S.[94] Other specimens, mostly from the same country, were received in subsequent years[6]. The examples of this animal just spoken of as being the first to arrive in England formed the subjects of a beautiful drawing by Mr. Wolf, a coloured lithograph taken from which has been published in the first volume of Wolf and Sclater’s ‘Zoological Sketches’ (plate xxii.).

Our figures of this species for the present work (Plate LV.) have been prepared by Mr. Smit from the descendants of the animals brought by Dr. Aitchison from Northern Persia, now living in the Society’s Gardens.

The series of specimens of this species in the British Museum comprises a skull from near Ispahan in Persia, presented by Dr. W. T. Blanford, F.R.S.; a head-skin and some horns from Gulran and Galicha, on the Afghan frontier, collected by Dr. Aitchison during the Afghan Boundary Commission; and some skins and skulls from the Saiar Mountains, Altai, presented by Mr. St. George Littledale. There are likewise a skin from the River Aksu, in Chinese Turkestan, presented by Major C. S. Cumberland, and several fine skulls and pairs of horns from the plains of Yarkand, obtained by the late Mr. Dalgleish, and presented to the Museum by Mr. A. C. Hume, C.B. All these last-named specimens represent the Yarkand subspecies, Gazella subgutturosa yarkandensis.

January, 1898.

THE BOOK OF ANTELOPES, PL. LVI.

J. Smit del. et lith.

Hanhart imp.

The Marica Gazelle.

GAZELLA MARICA.

Published by R. H. Porter.

[95]

86. THE MARICA GAZELLE.
GAZELLA MARICA, Thos.
[PLATE LVI.]

Closely allied to G. subgutturosa, with which it shares the substitution of white for the dark colour of the central facial band, the general plan of coloration, and the curvature of the horns. Size, however, very markedly smaller. General colour pale fawn. Facial markings almost obsolete; when distinguishable they are only of the general body-colour and very slightly defined from the paler bands between them. Ears long, their backs whitish fawn. Pale lateral band scarcely visible; dark lateral band and pygal band pale brown, little marked, scarcely darker than the dorsal colour. Limbs whitish throughout; distinct knee-tufts present.

Skull and horns, so far as the male is concerned, very much as in G. subgutturosa, although much smaller. Basal length of skull (in an old male) 6·1 inches, greatest breadth 3·15, muzzle to orbit 3·45.

Female. Similar, but horns only from 3 to 5 inches in length, slender, straight or slightly curved.

This little Gazelle is a recent discovery of Surgeon Lieut.-Col. A. S. G. Jayakar, C.M.Z.S., who has been for many years resident at Muscat in the service of the British Indian Government. Surgeon Jayakar, whose magnificent[96] collections of Muscat fishes are known to all ichthyologists, has during the past years collected and presented to the National Museum several consignments of mammals from this little-known country. Of these Thomas published an account in the ‘Proceedings’ of the Zoological Society for 1894, the most remarkable of them being a new Wild Goat, from the Akhdar Range behind Muscat, which was named Hemitragus jayakari after its discoverer. In 1897 the British Museum received from Surgeon Jayakar another consignment of mammals collected at Muscat within the previous two years. In this last series, besides the Oman specimens which were referable to species already recorded in Thomas’s paper, there were several skins and skulls of the present Gazelle, obtained from the Nejd or Nedsched Desert in the interior of Arabia. Thomas established his Gazella marica upon these examples.

In a letter addressed to Thomas, Dr. Jayakar says that four of the “Reem Gazelles” were from the Nejd Desert and one from Dahireh, the north-western district of Oman. “It is probable,” he continues, “that the species extends down to the desert behind Oman, as that is continuous with the Nejd Desert.” Surgeon Jayakar subsequently presented to the Museum a sixth (female) specimen from Aboor near Adam in Oman.

The Marica Gazelle is clearly a close relative of the Persian Gazelle, which it seems to represent in Arabia. But it is considerably smaller in size, paler in colour, and is nearly free from face-markings, besides having horns in the female sex. This last point is interesting, as it shows how little importance, in a generic sense, should be attributed to the presence or absence of horns in the female of an Antelope; for it appears that this species, in which the horns are present in the female, is unquestionably more nearly related to one in which the horns are absent in the female than to the group of Gazella dorcas, in which the horns are developed in both sexes.

In February, 1892, the Zoological Society of London received as a gift from Lt.-Col. Talbot, then British resident at Muscat, along with a Beatrix Antelope (Oryx beatrix), a small female Gazelle, with the information that it had been obtained from the Bahrein Islands, in the Persian Gulf. Sclater was at first much puzzled to give a name to this Gazelle, but after some hesitation came to the conclusion that it might be a small female of the Indian Gazella bennetti, which is known to extend along the coast of Baluchistan nearly to[97] the Persian Gulf, and accordingly entered it in the Society’s Register[7] under that name. This animal, however, which is still living in the Society’s Gardens, is undoubtedly a female of the present species. On reference to the late Theodore Bent’s paper on the Bahrein Islands (P. R. G. S. xii. p. 8, 1890) it will be found stated that on the desert which occupies the greater part of the largest island of the group “a small Gazelle is abundant,” and is often hunted by the Bahreini Arabs with hawk and hounds. There can be little doubt that this Gazelle is G. marica.

Our illustration of the Marica Gazelle (Plate LVI.) has been taken by Mr. Smit from the typical specimen from Nejd in the British Museum, and represents an adult male.

January, 1898.

[98]

THE BOOK OF ANTELOPES PL. LVII.

Wolf del J Smit lith

Hanhart imp.

The Dorcas Gazelle.

GAZELLA DORCAS.

Published by R. H. Porter

[99]

87. THE DORCAS GAZELLE.
GAZELLA DORCAS (Linn.).
[PLATE LVII.]

Size small; height at withers 21–22 inches. General colour pale fawn, rather variable in tone. Facial markings distinct; central band rufous fawn; streaks from eye to mouth brownish fawn, contrasting with the white bands between them. Ears long, whitish fawn behind. Light lateral band present, but not strongly marked; dark lateral band brown, considerably darker than the back, but not black. Pygal band indistinct. Knee-tufts present.

Skull rather lightly built; nasals narrow; anteorbital fossæ large and deep. Basal length 6·6 inches, greatest breadth 3·35, muzzle to orbit 3·55.

Horns of medium length, but quite different in their shape to those of any other species, although the difference is not very easy to explain. Primarily it may be said that they are flattened laterally, are evenly divergent as they curve backwards, reapproach terminally, and have their tips bent upwards in a well-marked curve. But in addition to this simple curvature, the middle portion of each horn is lyrated outwards, so that the longest diameter of the horn-section above is quite in a different plane to what it is at the base; it is in consequence of this lyration that the horns, apart altogether from the curved tips, reapproach each other terminally, while in all other species such reapproach as occurs is entirely due to the incurving of the tips. The lyration and curvature of the horns are well shown in our figure (p. 108), and a comparison of it with those of GG. przewalskii, marica, tilonura, and soemmerringi will show how different the method of terminal approximation is in this species as compared with them.

Female. Similar to the male, but horns slender, slightly curved, from one-half to three-fourths the length of those of the male.

[102]

Like other Antelopes known to Linnæus, the Dorcas Gazelle was placed by the great founder of systematic nomenclature in the genus Capra, and called Capra dorcas. The specific term selected was taken from the Greek, in which language it signifies primarily a wild goat or fawn, and subsequently the name of a woman, being, as we know from a familiar passage in the New Testament, the equivalent of the Syriac “Tabitha.” The term “dorcas,” however, according to good authorities, was also applied to the present animal by Ælian and other ancient writers. Linnæus based his Capra dorcas upon the Gazella africana of Ray’s ‘Synopsis of Quadrupeds,’ which there can be little doubt was intended for the present species, although it has been supposed by some authors to be rather applicable to the Bubal (Bubalis boselaphus).

In his memoir on the Antelopes, published at Berlin in 1767, Pallas placed Antilope dorcas eighth in the list, basing it on Buffon’s “La Gazelle,” which it is evident, both from the figure and the description, was taken from a specimen of the present animal.

Besides “La Gazelle” in the twelfth volume of his ‘Histoire Naturelle,’ Buffon also described and figured another Antelope, “Le Kevel,” of which he does not state the locality. Some authors have been disposed to refer Buffon’s Kevel to the larger Antelope of Algeria, which is generally called Gazella cuvieri. But Buffon’s description of the colour of the face and the length he attributes to the ears, as likewise his phrase that the Kevel is “plus petit que la Gazelle,” taken together form conclusive evidence against this view, and there can practically be no doubt that Buffon’s “Kevel” was a small individual of Gazella dorcas.

A third name invented by Buffon for one of the Gazelles, “La Corine,” has likewise been the source of some confusion. His figure and description were taken from a female animal at one time living in the park of Saint Cloud, but its locality was not given. Owing to the fact that some horns, brought home by Adanson from Senegal, were subsequently referred to Buffon’s “Corine,” the name Gazella corinna (founded upon Buffon’s “Corine”) has been sometimes applied to the Gazella rufifrons of Senegal. But, so far as we can judge from Buffon’s figure and description, the real[103] type of Buffon’s “Corine” must have been merely an ordinary female of Gazella dorcas, although there is some difficulty on the subject in consequence of the discrepancy between Buffon’s figure and his description. As Thomas has shown in his observations upon this point (P. Z. S. 1894, p. 469), it seems that a wholly satisfactory determination of Buffon’s “Corine” is barely possible.

It is manifest, however, that the names “kevella” and “corinna” cannot be safely assigned to any other Gazelle than Gazella dorcas.

The many systematists whose works we have quoted above in our list of synonyms added very little to our knowledge of this Gazelle beyond the fact that it was supposed to be distributed all over North Africa from Morocco to Egypt and to be also found in Palestine and Syria. It is only quite recently that we have ascertained some precise facts respecting the ranges of this and other Gazelles in the countries above mentioned, and even now our information on this subject is by no means perfect.

Commencing with Morocco, there is no doubt, we believe, from the testimony of various travellers, that a small Gazelle of this group does occur in the interior of that country and that it is probably of this species, but we have never had an opportunity of examining Moroccan specimens. In Algeria we have more definite information available.

From the days of Shaw the “Common Gazelle” has been recognized as an inhabitant of the “Barbary States.” The French naturalist Loche included it in his catalogue of 1858, but is not clear in distinguishing it from its allied species. Canon Tristram, in the ‘Great Sahara,’ published two years later, informs us that this Gazelle is found in small troops in every portion of the Sahara, and is the principal large game to be depended on for food, especially in the neighbourhood of the dayats beyond Laghouat, where pasturage is abundant. The fawns are dropped in the early summer, and follow the dam until towards the end of the autumn. The Bedouin gather the droppings, which have a strong aromatic scent, to mix with snuff.

In the pages of the ‘Exploration Scientifique de l’Algérie’ devoted to Mammals, likewise from the pen of Loche, we find a little more information concerning the Algerian Gazelles, but it does not appear that Loche was at all clear in discriminating the various species that are there met with.

Good and precise information has, however, been given us on this subject by Mr. Alfred E. Pease, M.P., in his article on the “Antelopes of the Aures[104] and Eastern Algerian Sahara,” published in the Zoological Society’s ‘Proceedings’ for 1896, the principal portion of which we must quote at full length:—

Passing on to the Beylik of Tunis, we have excellent notes on the Gazelles of this country drawn up by Mr. Joseph I. S. Whitaker, F.Z.S., published in the same volume of the Zoological Society’s ‘Proceedings.’ Mr. Whitaker writes of this Gazelle as follows:—

So little is known of the natural history of Tripoli and Barca that we can only presume that the Dorcas Gazelle ranges through these countries on its way to Egypt, where it is well known to be abundant in the Western Desert. Sclater examined large numbers of both sexes of this species in the Zoological Garden of Gizeh near Cairo in 1895 from this locality[8], and several specimens from the same source have been received in exchange by the Zoological Society of London. Our figures of both sexes of the Dorcas Gazelle (Plate LVII.) have been prepared by Mr. Smit from examples thus obtained.

In the eastern desert of Egypt the Dorcas Gazelle appears in these days to be not nearly so common. Mr. E. N. Buxton, who traversed the eastern desert in his expedition after Capra sinaitica, tells us that two or three Gazelles together were the most he ever saw at one time. Between the Nile and the granite mountains 80 miles to the east, a very arid district, Mr. Buxton only saw Gazelles once. They were more numerous among the foot hills of the Kettar range and the porphyry mountains, for the obvious reason that there is more vegetation there.

The Gazelles frequently depicted in the paintings of the ancient Egyptian tombs and temples were, no doubt, usually Gazella dorcas in Lower, and G. isabella in Upper Egypt, although they were probably also well acquainted with G. arabica. Dr. Hartmann in his interesting disquisition on the animals of these paintings (Zeitschr. f. Aegyptische Sprache und Alterthumskunde,[106] 1864, p. 22) gives us the hieroglyphic symbols of the Gazelle, and its corresponding name as “Gahés.” It was evidently a common object of chase even in those days.

Crossing over into the Holy Land, we find the Dorcas Gazelle registered in Canon Tristram’s ‘Fauna and Flora of Palestine’ as met with in all suitable localities. From the same author’s ‘Natural History of the Bible’ we extract the following passages relating to this favourite animal:—

Canon Tristram describes the mode of hunting Gazelles practised by the Arabs as follows:—

In the desert country east of the Jordan, Canon Tristram tells us, the Dorcas Gazelle is replaced by the Arabian Gazelle (G. arabica); but a Gazelle, probably of this species, is found in the Syrian Desert north of Damascus, as testified by many writers.

In his interesting volume on ‘Palmyra and Zenobia,’ Dr. William Wright, describing his journey between Damascus and Palmyra, says:—

But, as we are informed in the valuable papers on the Mammals of Asia Minor published by Messrs. Danford and Alston in the Zoological Society’s ‘Proceedings’ for 1877 and 1880, the Dorcas Gazelle ranges far north of Syria. Danford states that it is “not uncommon” on the plain of Tchukurova and about Tarsus and Adana in the south-east of Asia Minor, and that it is “common” in the wooded valley of the Pyramus on the plain of Bazardjik and extends thence into the stony wooded uplands on the right bank of the Northern Euphrates.

When taken young, the Dorcas Gazelle is easily tamed and becomes very docile and affectionate. It is frequently kept in captivity by the Arabs and thus passes into the hands of Europeans who visit the East. As will be seen by reference to the Zoological Society’s List of Animals, specimens of this species reach the Gardens every year. But they cannot be said to thrive in the climate of England, where they miss the bright sun and dry air of their native deserts, and seldom produce young.

The series of examples of this Gazelle in the National Collection is by no means a full one, and wild-killed examples with ascertained localities from different parts of its range are much wanted. At the present[108] time, besides a number of old specimens without localities, there are only in the collection an adult male from Biskra in Algeria presented by Sir Edmund Loder, a pair of skins from the same place presented by Messrs. Rowland Ward and Co., and a skull from Egypt presented by the late Sir Gardner Wilkinson. The accompanying illustration (fig. 57) gives a front view of a good head of this Gazelle prepared by Mr. Smit from these specimens.

January, 1898.

Fig. 57.

Head of the Dorcas Gazelle, ♂.

(From specimens in the British Museum.)

THE BOOK OF ANTELOPES, PL. LVIII.

J. Smit del. et lith.

Hanhart imp.

The Edmi Gazelle.

GAZELLA CUVIERI.

Published by R H Porter.

[109]

88. THE EDMI GAZELLE.
GAZELLA CUVIERI (Ogilby).
[PLATE LVIII.]

Size comparatively large; height at withers about 26 or 27 inches. Hair rather long, rough and coarse. General colour dull fawn. Face-markings distinct; the central facial band brownish fawn, with a black patch on the top of the nose, in front of which the muzzle is white. Ears long, pointed, their backs fawn. Dark lateral line and pygal band distinct, darker than back; light lateral line present, but little defined. Knee-brushes distinct.

Basal length of skull 7·35 inches, greatest breadth 3·6, muzzle to orbit 4·45.

Horns rather short in proportion to the size of the animal, thick, strongly[110] ribbed, very slightly curved backwards, and but little divergent from each other; the tips slightly curved upwards and forwards.

Female. Similar, but horns shorter, slenderer, and straighter.

The “Edmi” or Mountain Gazelle of Algeria, though it has often been confounded with the Dorcas, and has only been accurately known within the last few years, is without doubt an absolutely different species not only in structure, but in habits and mode of life. As Sir Victor Brooke has pointed out, it is easily distinguished from all its allies by its larger size, rough coat, dark colour, and long ears.

The first published information that we can certainly refer to this species is that of Frédéric Cuvier, who figured both sexes in his folio work on Mammals from specimens living in the Jardin des Plantes at Paris, but called it only “le Kevel gris,” without giving it any scientific designation.

Some time in 1839 a living female example of this Antelope was presented to the Zoological Society of London by Mr. W. Willshire, one of their corresponding members, who had procured it at Mogador. After its death in May 1840, Mr. Ogilby, who was at that time Secretary of the Society, and was specially interested in the study of mammals, brought the specimen before the notice of the Zoological Society at one of their scientific meetings, and proposed to name the species “cuvieri” after M. Frédéric Cuvier. Ogilby stated that he had observed examples of the same Gazelle in the Paris Museum, and that M. Cuvier would have described it had he, Ogilby, not done so. There can be no question therefore of Ogilby’s animal being the same as Cuvier’s “Kevel gris,” and that Gazella cuvieri is the earliest certain name to adopt for it.

In 1849 Fraser published a figure of this species in his ‘Zoologia Typica,’ taken from Ogilby’s typical specimen, which is now in the British Museum. Although imported from Mogador there can be little doubt that this example was originally obtained from the chain of the neighbouring Atlas. The Gazelles observed in the Great Atlas in company with Wild Sheep (Ovis tragelaphus) by Mr. W. B. Harris, F.R.G.S., on his journey from Morocco to Tafilat in 1893, were no doubt Gazella cuvieri.

Passing on to Algeria we find that Loche appears to have referred to this[111] Antelope under the name Gazella corinna, and that Canon Tristram and Lataste have called it Gazella kevella. We have already shown, however, that both these names are properly to be applied to the Dorcas Gazelle.

In 1890, Mr. Edward Buxton met with this Gazelle during a shooting excursion into the Atlas, of which he has given us a most interesting account in one of the chapters of his ‘Short Stalks.’ Mr. Buxton’s principal object of pursuit on that occasion was the Aroui or Barbary Sheep (Ovis tragelaphus), but he also had the good fortune to obtain a fine head of the Mountain Gazelle, which he exhibited at a meeting of the Zoological Society on March 31st of that year (see P. Z. S. 1890, p. 363).

Mr. Buxton tells us, in the course of the remarks he made on this occasion, that the Mountain Gazelle of Algeria is “about twice the size of the Gazelle of the plains (Gazella dorcas), and has straight instead of lyre-shaped horns. It lives on the same kind of steep ground as the Aroui, perhaps at a rather lower elevation. The fact that it is essentially a mountain animal is, I think, shown by its large callous knees, like those of a London cab-horse. The Aroui has the same. They are, I think, absent in Gazella dorcas. Another feature consists in the curious hollows or pouches on each side of the testicles.”

In his ‘Short Stalks’ Mr. Buxton gives us full particulars of his adventures in obtaining the much-coveted head of this animal above referred to, and illustrates them by a beautiful picture of a group of these Gazelles drawn and engraved after his instructions by Mr. G. E. Lodge.

In his field-notes on the Antelopes of Eastern Algeria, published in the Zoological Society’s ‘Proceedings’ for 1896, Mr. A. E. Pease speaks as follows of the present species:—

[112]

Crossing the frontier of Algeria into Tunis we find the Edmi Gazelle prevalent in suitable districts throughout that country. Mr. J. I. S. Whitaker, F.Z.S., who knows Tunis and its birds and mammals well, writes, in the same volume of the ‘Proceedings’ as we have quoted above, of his experiences of this animal as follows:—

Among the wood-blocks left ready for use by the late Sir Victor Brooke[113] was a figure of a fine head of a male of this Gazelle drawn by Mr. Smit, of which we now give an impression.

Fig. 58.

Head of Edmi Gazelle, ♂.

(Drawn by Smit under the direction of
the late Sir Victor Brooke.)

The Edmi Gazelle is not often brought alive to Europe, but besides Ogilby’s type specimen, which we have already mentioned, at least three others have been exhibited at various times in the Zoological Society’s menagerie. An example of this species was first obtained in 1839, as mentioned in the Report of the Council for 1840, and another was acquired in November 1862. Others were presented by Capt. Alan Gardner, R.N., in June 1865, and by Rear-Admiral Sir William Hall, R.N., in May 1867. Sclater observed a female of this Gazelle in the Zoological Garden of Berlin in September 1897 (see P. Z. S. 1897, p. 813).

As we have already stated, the typical specimen of this Gazelle, formerly in the Zoological Society’s Collection, is now in the British Museum, as is also a stuffed female, originally presented by H.M. the Queen to the Zoological Society, but transferred to the National Collection in 1855. In the[114] British Museum Gallery of Mammals will be found a good adult stuffed specimen of this Gazelle stated to be from near Biskra, Algeria, and presented by Mr. J. I. S. Whitaker. There are also some frontlets and horns of this species from the same locality presented by Messrs. Rowland Ward & Co.

We are greatly indebted to Sir Edmund Loder for a photograph of a head of a female of this Gazelle, taken from a specimen in his collection, which he himself shot on the 27th of February, 1893, on the Ahmar Khaddou Mountains, two days’ march east of Biskra. It shows very clearly the inferior size of the horns in this sex, and the long ears characteristic of the species.

Fig. 59.

Front view of head of Edmi Gazelle, ♀.

(From a photograph.)

Our illustration of the male of this Gazelle (Plate LVIII.) has been drawn by Mr. Smit from the Algerian specimen in the British Museum above referred to.

January, 1898.

THE BOOK OF ANTELOPES, PL. LIX.

J. Smit del. et lith.

Hanhart imp.

The Arabian Gazelle.

GAZELLA ARABICA.

Published by R. H Porter.

[115]

89. THE ARABIAN GAZELLE.
GAZELLA ARABICA (Licht.).
[PLATE LIX.]

[116]

Size medium; height at withers about 24 or 25 inches. General colour dark smoky fawn, much darker than in the allied species. Facial markings distinct; central facial band dark rufous-fawn, with a black spot on the nose. Ears of medium length, brownish-fawn behind. Dark lateral and pygal bands smoky brown; light lateral band very slightly lighter than the back. Limbs more rufous than body; knee-brushes brown or black.

Skull with short broad nasals; anteorbital fossæ shallow in the only good skull available. Basal length 6·75 inches, greatest breadth 3·3, muzzle to orbit 3·7.

Horns thick and rather short, almost straight and parallel to each other, a little curved backwards below, and forwards above.

Female. Similar to the male, but horns short and straight.

Although the Arabian Gazelle was described and figured as long ago as 1827, and specimens of it are by no means rare in captivity, we have as yet received little information about its exact range and its mode of life. But the great peninsula of Arabia still remains, we must recollect, one of the largest tracts on the earth’s surface that has been least explored by scientific travellers.

Hemprich and Ehrenberg, the discoverers of this Gazelle, met with it during their travels on the eastern coast of the Red Sea, and transmitted specimens of both sexes to the Berlin Museum. Here they were first described and figured by Lichtenstein in his ‘Darstellung der Säugethiere’—a work devoted to making known the riches of the Mammal-collection of the great Institution of which he was the Director.

Following Hemprich and Ehrenberg’s MS., Lichtenstein named the species “Antilope arabica,” and a short time afterwards it was again described and figured by Ehrenberg in his ‘Symbolæ Physicæ’ under the same designation.

Ehrenberg informs us that he and his fellow-traveller Hemprich obtained their first specimens of this Gazelle at Hamam el Faraun, on the coast of the Sinaitic Peninsula between Suez and Tor, and subsequently found it abundant on the island of Farsan on the Arabian coast of the Red Sea. They also observed Gazelles which they believed to be of the same species near Baalbec in Syria, but these, we think, are more likely to have been Gazella dorcas.

[117]

Succeeding authorities have added very little to our knowledge of this Gazelle. Canon Tristram, in his ‘Fauna and Flora’ of Palestine, mentions a Gazelle occurring in the “desert-country east of the Jordan” as being probably of this species; but we believe that he did not obtain any good specimens of it. Dr. Blanford, in his volume on the ‘Geology and Zoology of Abyssinia,’ has figured (for comparison) a head of this species obtained by Captain Heysham near Mocha, S.W. Arabia; and in the Zoological Society’s ‘Proceedings’ for 1874, the late Sir Victor Brooke gave a woodcut of the head of this Gazelle, which, by the kind permission of the Society, we are enabled to reproduce.

Fig. 60.

Head of Arabian Gazelle.

(P. Z. S. 1874, p. 141.)

Living examples of the Arabian Gazelle are easily obtained at Aden and at Hodeidah, Jeddah, and other Arabian ports on the Red Sea, and are often brought to Europe. We have little doubt that the Gazelles in the Derby Menagerie figured by Waterhouse Hawkins in the third plate of the ‘Gleanings,’ and there called by Lord Derby’s MS. name, Gazella vera, were of this species, though in the text they are referred to as G. dorcas and in[118] the list of plates as G. cuvieri. The Zoological Society of London appear to have received their first specimens in 1874[9], and since that date (as will be seen by their published Lists of Animals) have acquired many examples, chiefly by presentation. At the present time there are two fine males in the Society’s Gardens, both brought from Aden and presented—one by Mr. R. G. Buchanan and the other by Mr. J. Benett Stanford, F.Z.S. From the former of these Mr. Smit’s drawing (Plate LIX.) was taken.

The British Museum has lately acquired from the Zoological Society a good male example of this Gazelle, which was obtained at Aden and brought alive to London. It has been mounted for the Mammal Gallery. The Museum also has the skull from Mocha figured by Dr. Blanford, as already mentioned, and since presented by him to the collection, and the skin of a young animal from Gilead, obtained by Canon Tristram, the determination of which is, however, somewhat doubtful.

January, 1898.

THE BOOK OF ANTELOPES, PL. LX.

J. Smit del. et lith.

Hanhart imp.

The Indian Gazelle.

GAZELLA BENNETTI.

Published by R. H. Porter.

[119]

90. THE INDIAN GAZELLE.
GAZELLA BENNETTI (Sykes).
[PLATE LX.]

Size medium; height at withers 24 to 25 inches. General colour dull fawn. Facial markings distinct, the darker ones rufous-fawn; a black spot on the top of the nose. Ears of medium length, fawn-coloured behind. Dark lateral and pygal bands brownish fawn, scarcely darker than the back; light lateral bands scarcely perceptible; knee-brushes present.

Skull with deep anteorbital fossæ:—Basal length 7·2 inches, greatest breadth 3·45, muzzle to orbit 4.

Horns thick, heavily ribbed, close together, diverging little but evenly, gently curved backwards below and forwards at their tips.

Female. Similar to the male, but horns straight, simple, about two-thirds the length of those of the male.

Like the Lion and the Cheetah this Gazelle belongs to an Ethiopian type of mammals, and was originally, no doubt, an intruder into India from the west. But, as will be seen when we come to describe its range, it has now spread itself over the greater part of the peninsula except on the eastern[121] side. On the west the Indian Gazelle extends far along the Mekran coast to the Persian Gulf, and there meets the Arabian Gazelle, of which it is undoubtedly a very close ally, although the latter is always much darker on the back.

Although the Indian Gazelle, or “Ravine-Deer,” as it is usually termed by Europeans, was doubtless known to the sportsmen of British India long ago, it was not made known to science until 1831, when Col. Sykes, one of the earliest pioneers in Indian natural history, described it in a communication made to the Zoological Society of London. Sykes, in his paper on the Mammals of the Deccan read before the Society in July of that year, proposed to name it Antilope bennetti, after the late Edward Turner Bennett, a well-known naturalist, who was at that time Vice-Secretary of the Society. Sykes met with this Antelope on the rocky hills of the Deccan “in groups rarely exceeding three or four in number, and very frequently solitary.” In 1849, Fraser published a figure of this species in his ‘Zoologia Typica,’ taken from one of Sykes’s male specimens in the British Museum, which is still in the National Collection, although not in the exhibition gallery. In 1844, in his description of the mammals of Jacquemont’s ‘Voyage dans l’Inde,’ Isidore Geoffr. St. Hilaire described and figured an Antilope hazenna, which he at that time considered to be different from the present animal. But there can be no doubt that Jacquemont’s specimens, which were obtained at Malwa in Central India, are the same as Gazella bennetti, and Sykes’s term being the oldest has been universally employed as the designation of this species. As we shall presently show, Gazella christii of Gray, from Sind, and Gazella fuscifrons of Blanford, from Baluchistan, are names which have been based on what are merely slightly divergent forms of Gazella bennetti.

From the researches of Elliot, Jerdon, Blyth, Blanford, and other authorities on the mammals of British India, we are now well acquainted with the range of this Gazelle in the peninsula and adjoining lands to the west. Dr. Blanford describes it as extending throughout the plains and low hills of North-western and Central India, and thence through Baluchistan to the eastern shore of the Persian Gulf. In the Indian peninsula, he continues, the Indian Gazelle ranges in suitable localities throughout the Punjab, Sind, Rajputana, the N.W. Provinces, and the whole of the Bombay Presidency with the exception of the Western Ghats and Konkan; it also occurs in Central India as far east as Palamow and Western Sargiya, and in the[122] Central Provinces as far east as Seoni and Chánda, together with the Hyderabad territories and the Madras Presidency to a little south of the Kistna, Gazelles being found at Anantapur, south of Kurnool, and in Northern Mysore.

For an account of the habits of the Indian Gazelle and the modes of its chase, we cannot do better than refer to the last edition of General Kinloch’s ‘Large Game Shooting,’ where they are described as follows:—

Dr. Blanford tells us that this Gazelle lives on grass and on the leaves of bushes, and, so far as he is aware, never drinks. “I have seen it,” he says, “in the deserts of Sind in places where the only water for twenty miles round was procured from wells; and in spots in Western and Central India where, in the hot weather, the only water to be obtained was in small pools remaining in the beds of streams. But around these pools, in which the tracks of almost every animal in the forest was to be seen, I never yet saw the very[123] peculiarly formed tracks of the Gazelle, although it frequently abounded in the neighbourhood. The four-horned Antelope, on the other hand, drinks habitually.”

Gazella christii, which we have alluded to above as synonymous with this species, was a MS. name of the late Dr. Gray, which appears to have been first published by Blyth in 1842, and applied to a pale form of the present animal from Kutch and Sind. But more recent researches have shown that it is not properly separable from the typical Gazella bennetti.

Gazella fuscifrons, another synonym also mentioned above, was based by Dr. Blanford in 1873 on a doe with distinctly ringed horns and with portions of the face dark brown, obtained in Baluchistan. But the late Sir O. B. St. John subsequently procured what he justly concluded to be the male of this form, which, as acknowledged by Dr. Blanford himself, proved to be not distinct from Gazella bennetti. By the kindness of the Zoological Society we are enabled to reproduce Dr. Blanford’s figures of the head of Gazella fuscifrons, which, except for the slight differences above mentioned, give an equally good idea of the head of the typical G. bennetti.

Fig. 61.

Head of Gazella fuscifrons, ♀.

(P. Z. S. 1873, p. 317.)

[124]

The Indian Gazelle is frequently brought to Europe alive, though it is not so common in our menageries as Gazella dorcas, G. subgutturosa, and some other species. According to the Zoological Society’s books, the first examples were received in 1838, and since 1860, as will be seen by the printed lists, about twelve specimens have been exhibited. A pair presented by Capt. H. J. Hope Edwards, in April 1883, bred, and the female gave birth to a young one in November of that year; but, like other Gazelles, this species does not usually thrive in the dull climate of England.

As is the case with many common animals, the British Museum does not contain a good series of this Gazelle, and specimens with exact localities from all parts of its range are much required. Besides the old mounted examples from the Deccan presented by Col. Sykes and already alluded to, it possesses only a skin from Sind received from the Karachi Museum, and several skulls and pairs of horns from the Salt Range of the Punjâb and Kelat, received in the Hume Collection.

Our figures of this species (Plate LX.) have been prepared by Mr. Smit—the male from the skin received from the Karachi Museum, and the female from Col. Sykes’s specimen.

January, 1898.

THE BOOK OF ANTELOPES, PL. LXI.

J. Smit lith.

Hanhart imp.

Speke’s Gazelle.

GAZELLA SPEKEI.

Published by R. H. Porter.

[125]

91. SPEKE’S GAZELLE.
GAZELLA SPEKEI, Blyth.
[PLATE LXI.]

Size slightly greater than in G. dorcas; height at withers 23–24 inches. Body pale brownish fawn, the light lateral band but little paler than the back, then dark or dull blackish, not sharply defined. Central facial band brownish fawn; top of muzzle, over nasal bones, with a distinct blackish patch, in front of which there is a peculiar swollen and corrugated cushion of skin raised up above the level of the face, and extensible at the pleasure of the animal. Dark cheek-band narrow, indistinct, the light band above it broad and extending to the muzzle. Ears long,[126] narrow, pointed, their backs whitish fawn. Knee-tufts present, brownish fawn. Pygal band very indistinct.

Skull with short broad nasals, the premaxillæ not or barely touching their outer corners. Basal length in an old male 6·5 inches, greatest breadth 3·35, muzzle to orbit 3·6.

Horns but slightly divergent, evenly and strongly curved backwards for three-fourths their length, their tips gently recurved upwards.

Female. Like the male, but the horns slender, little ridged, less curved, about three-fourths the length of those of the male.

There can be no doubt that the two Gazelles which inhabit the maritime plain and the high inland plateau of Somaliland respectively, although they are closely allied, and have been confused together by some writers, belong to distinct species, distinguishable by well-marked characters. The Gazelle of the interior plateau, which we treat of first, when compared with that of the coast-land is at once recognizable by the generally browner colour, the darker lateral band, the black nose-spot, and above all by the wrinkled and elevated nose of the adult, which is not met with in the sister species.

Speke’s Gazelle was first discovered by the energetic African explorer, whose name it appropriately bears, during his expedition to Harar in the summer of 1854 in company with the late Capt. Sir Richard Burton[10]. Speke, who attended to the natural history of the expedition, forwarded the collections made upon this occasion to Blyth, at that time curator of the Asiatic Society’s Museum at Calcutta, and in the zenith of his zoological work. In his report upon the collection, which was published in the twenty-fourth volume of the ‘Journal of the Asiatic Society of Bengal,’ Blyth did not venture to bestow a new name on this Gazelle, although he gave an accurate description of it, and added a note (obtained from Burton) calling special attention to “the elevation of loose replicated skin upon the nose,” so that there can be no doubt as to which of the two allied species Speke’s specimens (which are still in the Calcutta Museum[11]) belong.

[127]

In the reprint of Blyth’s ‘Report,’ which was edited by Speke in 1860 after his return to this country, this Gazelle was erroneously referred to G. cuvieri of Ogilby. In 1883, however, Blyth, who had discovered that this was a mistake, proposed the name Gazella spekei for this species in his ‘Catalogue of the Mammals of the Asiatic Society’s Museum,’ and this appellation has been generally adopted for it ever since.

When the late Sir Victor Brooke wrote his Monograph of the Gazelles in 1873 Speke’s Gazelle was hardly known in this country, and Brooke was only acquainted with it from photographs of the type specimens in the Calcutta Museum. But since that date Somaliland has been fully opened to British travellers, and the numerous explorers and sportsmen who have visited that much-hunted country have brought back good sets of specimens both of Speke’s and of Pelzeln’s Gazelle, and made us well acquainted with the ranges and other peculiarities of these two species.

One of the first British travellers who visited Somaliland, and made the acquaintance of Speke’s Gazelle, was the late Mr. F. L. James, who proceeded there on a shooting-expedition in January 1884, accompanied by his brother and Mr. E. Lort Phillips[12]. Mr. Lort Phillips read some notes on the Antelopes obtained on this occasion before the Zoological Society in December 1885, and in alluding to this Gazelle called it the “Flabby-nosed Gazelle,” to which term Sclater attached a footnote stating that it was “probably of a new species,” but required further examination. This examination Sclater bestowed upon Mr. Lort Phillips’s specimens shortly afterwards (see P. Z. S. 1886, p. 504), and came to the correct conclusion that the so-called Flabby-nosed Gazelle was quite distinct from the species of the coast land. He unfortunately did not perceive that it was the species of the high plateau and not that of the coast land, which had already been named Gazella spekei by Blyth, and therefore gave it a new name, Gazella naso, under which appellation it will be found described and its characteristic head figured in Sclater’s article in the Zoological Society’s ‘Proceedings’ for 1886. But, as Thomas has subsequently shown (P. Z. S. 1891, p. 210), there can be no doubt that Gazella naso is merely a synonym of Gazella spekei.

Another well-known author, who must not fail to be quoted in any[128] reference to the game animals of Somaliland, is Capt. H. G. C. Swayne, R.E. Capt. Swayne has made no less than seventeen trips to that attractive country, and is probably better acquainted with its larger mammals than any other living individual. In his excellent narrative of his adventures[13], Capt. Swayne has given us some capital notes on Speke’s Gazelle and its near ally Pelzeln’s Gazelle, both of which are known to the natives by the same name “Dhero.”

Fig. 62.

Head of adult male Speke’s Gazelle.

(Brit. Mus.)

Capt. Swayne calls the former Antelope the “Ogo, or Plateau Gazelle,” and the latter the “Guban, or Lowland Gazelle,” and describes the peculiarities of the present species as follows:—

Fig. 63.

Head of adult female Speke’s Gazelle.

(Mr. F. Gillett, F.Z.S.)

A still more recent explorer of Somaliland, Mr. A. E. Pease, M.P., F.Z.S., has most kindly favoured us with some excellent notes on Speke’s Gazelle and its sister species, which we cannot do better than reproduce. Mr. Pease has also sent us along with his MS. remarks a sketch-map of the northern part of Somaliland, in which the ranges of these two species are accurately shown.

[130]

He writes as follows:—

So far as we know, but one specimen of Speke’s Gazelle has as yet reached this country alive. This was a young male, presented to the Zoological Society’s Menagerie in November last year by Dr. L. de Gébert, who had obtained it at Djibutil, the French port of Abyssinia. Unfortunately it did[131] not live long in captivity, but after its death Sclater, with Mr. F. E. Beddard’s kind assistance, was able to examine the specimen more closely. It exhibited a slight protuberance on the nose, as shown in the figure (fig. 64), which by the kindness of the Zoological Society we are enabled to reproduce on the present occasion. Underneath the skin of the nose was a slight cavity, which was easily inflated into a protuberance by blowing air into the nostrils. But dissection, which was carried out by Mr. Beddard, revealed no trace of any glandular structure.

Fig. 64.

Head of young male Speke’s Gazelle.

(P. Z. S. 1897, p. 920.)

The collection of the British Museum contains a good mounted example of this Gazelle (procured from Herr Menges, and originally obtained from Gerbatir in Somaliland), from which the coloured figure (Plate LXI.) has been prepared by Mr. Smit. There is likewise in the Museum a good series of skins and skulls from different places in Somaliland collected and presented by Mr. T. W. H. Clarke, Capt. H. G. C. Swayne, R.E., Col. Arthur Paget, and Mr. Ford G. Barclay. From one of these our figure (p. 128) of a good adult head of this Gazelle has been prepared by Mr. Smit; while that of the female (p. 129) has been drawn from a mounted head kindly lent to us for that purpose by Mr. Frederick Gillett, F.Z.S.

May, 1898.

[132]

THE BOOK OF ANTELOPES, PL. LXII.

J. Smit lith.

Hanhart imp.

Pelzeln’s Gazelle.

GAZELLA PELZELNI.

Published by R. H. Porter.

[133]

92. PELZELN’S GAZELLE.
GAZELLA PELZELNI, Kohl.
[PLATE LXII.]

Size rather greater than in G. spekei; height at withers about 25 inches. Colour brownish fawn, rather more rufous than G. spekei. Light lateral band distinct; dark band rufous brown, similar to that of the back but rather darker in tone, not blackish. Pygal band distinct, brown. Centre of face dark fawn, without either the black spot or the pale swollen cushion characteristic of G. spekei. Dark and light cheek-bands short and indistinct. Knee-tufts dark brown.

Skull narrower than in G. spekei, and with long narrow nasals, which articulate broadly with the premaxillæ. Basal length in an old male 6·87 inches, greatest breadth 3·25, muzzle to orbit 3·75.

Horns more evenly divergent, much straighter and less curved backwards than in G. spekei, but otherwise similar. In length they attain to about 11 or 12 inches.

Female. Like the male, but the horns much smoother and slenderer, and only about three-fourths the length.

[134]

As we have already mentioned in our account of the last species, the late Mr. F. L. James and his party, who visited Somaliland in 1884, appear to have been the first to bring to England examples of the two allied Gazelles of Somaliland. Unfortunately, however, though perhaps not unnaturally, Sclater, who assisted Mr. E. Lort Phillips in the determination of the Mammals obtained during that expedition, referred the coast-land specimens to Gazella spekei, and described the examples from the high plateau as belonging to a new species, Gazella naso. About two years later, however, this error was corrected by Herr H. F. Kohl, of the Natural History Museum of Vienna, who, in an article upon new and rare Antelopes collected by Herr L. Menges in Somaliland, among which examples of both these Gazelles were comprised, rightly referred the upland species to Gazella spekei of Blyth and gave to the lowland species, then still unnamed, the title of Gazella pelzelni, after the late August von Pelzeln, a well-known naturalist, who was at that time Custos of the Imperial Museum of Natural History.

Thomas, in his article on the Antelopes collected in Somaliland by Mr. T. W. H. Clarke, published in the Zoological Society’s ‘Proceedings’ for 1891, was the first to make this matter perfectly clear, and to establish the name Gazella pelzelni as the permanent designation of the coast-land Gazelle of Somaliland. Since that date the distinctions between the two allied species have become well recognized and understood, and numerous examples of both species have been obtained by the naturalists and sportsmen who have recently visited that country.

Capt. Swayne, in his well-known work on Somaliland and its wild animals (from which, by the kind permission of Messrs. Rowland Ward & Co., the publishers, we have been allowed to borrow illustrations of the heads of both sexes of this Gazelle), tells us that the “short-coated, light-coloured Lowland Gazelle” carries rather longer horns than those of the Plateau Gazelle (Gazella spekei), which are “shorter, thicker, more curved, and better annulated.” “The habits of both,” he continues, “are nearly alike; they go in moderate-sized herds of from three to ten, and resort mostly to stony or sandy undulating ground or ravines thinly dotted over with mimosas. Both species are fond of salt and do not require water. It is hard to understand what they can pick up to eat in the wretched ground which they frequent. They have a curiosity which amounts almost to impudence, but are[135] wonderfully on the alert, and hard to shoot, seeming to know perfectly well the range of a rifle, and presenting but a very small target.”

Capt. R. Light, writing to Sclater in 1892, tells us that when he visited Somaliland in 1891 he found this Gazelle between Berbera and Zeila, close down by the sea: “they were often observed feeding side by side with camels and flocks of sheep and goats. When startled they move off the ground in a quick trot, taking bounds over any obstacles and finally breaking into a gallop.”

Fig. 65.

Head of Pelzeln’s Gazelle, ♂.

(From Swayne’s ‘Somaliland,’ p. 317.)

Fig. 65a.

Skull of Pelzeln’s Gazelle, ♀.

(From Swayne’s ‘Somaliland,’ p. 317.)

Mr. Pease, who has kindly supplied us with notes on this Gazelle as well as on the preceding species, writes that Pelzeln’s Gazelle is essentially the species of the maritime plain and could be seen within shot of the town of Bulhar when he was there in 1896, and within a mile or so of Berbera. “In life it appears of a light rich yellow-buff in colour, with the usual Gazelline marks rather faintly indicated. Its coat is short and fine, and its horns are[136] straighter and longer than those of G. spekei. The horns of the female are weak and almost smooth, like those of Speke’s Gazelle.” “Within fifty miles of the sea-shore,” he continues, “this Gazelle is exceedingly numerous in suitable places. Half-a-dozen herds may be often seen at a time, but I have seldom observed more than twenty in a single band. In size G. pelzelni is larger than G. spekei, the average height being about 25 inches, while the weight of the carcase is usually a little over 40 lbs.”

Mr. D. G. Elliot, who made a successful expedition to Somaliland in 1896, for the purpose of obtaining specimens for the Field Columbian Museum of Chicago, gives us the following account of his experiences with the present species:—

The British Museum contains a good male specimen of Pelzeln’s Gazelle, mounted from a skin obtained by Herr Menges near Berbera in Somaliland. Mr. Smit’s figures of this species (Plate LXII.), which represent the male in two positions, have been prepared from it. The Museum also contains two skins from the plains of Berbera, collected by Capt. Swayne and originally sent home to Sclater.

May, 1898.

THE BOOK OF ANTELOPES, PL. LXIII.

J. Smit lith.

Hanhart imp.

Loder’s Gazelle.

GAZELLA LEPTOCEROS.

Published by R. H. Porter.

[137]

93. LODER’S GAZELLE.
GAZELLA LEPTOCEROS (F. Cuv.).
[PLATE LXIII.]

Height of male at withers about 25 inches. General colour very pale sandy fawn, the Gazelline markings little defined. Central facial band and darker cheek-bands sandy, not rufous, and but little contrasting with the light facial streaks. Light lateral bands scarcely perceptible, and the darker ones below them only pale sandy with a tinge of brownish, as are the pygal bands, neither being much darker than the general dorsal colour. Ears long, narrow,[138] pointed, pale whitish buff externally. Tail sandy at base, darkening terminally to brownish black. Front of fore limbs sandy, of hind limbs whitish; knee-brushes distinct, but little darker than the general colour. Hoofs variable in shape, those of specimens from the sandy regions of the Sahara much elongated, while in other regions they are of the usual shape.

Skull of normal proportions; premaxillæ broadly articulating with nasals. Basal length in an old male 6·45 inches, greatest breadth 3·3, muzzle to orbit 3·6.

Horns of male long, about twice the length of the skull, slender, closely and heavily ringed nearly to the tip. They are very variable as to their exact curvature, but are ordinarily rather straighter than in other species, curving but slightly backwards; they are near together basally, diverging above, sometimes very widely, so as to make them resemble divergent horns of G. granti in miniature.

Female. Similar to the male, but the horns, although nearly equally long, are much slenderer and even less curved than in the male.

The great folio work of Geoffrey St.-Hilaire and Frédéric Cuvier entitled ‘Histoire Naturelle des Mammifères,’ which was issued in livraisons from 1824 to 1842, contains a long series of coloured figures of mammals, mostly taken from examples living in the well-known Menagerie attached to the Jardin des Plantes. Amongst these in the seventy-second livraison, published in 1842, were the first descriptions and figures given of both sexes of the present Gazelle, from examples stated to have been brought from Sennaar by Burton. They had lived in the Menagerie, we are told, two years, and had bred a young one, which resembled its parents in most particulars. The appropriate scientific name “leptoceros,” from the long thin horns, had been given to this species, we are informed, by Georges Cuvier, and was adopted by the authors of the work referred to, who, however, called it at the head of their article, after their usual fashion, only by the French name “Antilope à longues cornes.”

Very little more information was acquired concerning this Gazelle for many years. Most of the systematists were entirely unacquainted with it,[139] and could only quote the original descriptions. Sundevall and Gray considered it to be merely a variety of Gazella dorcas. Rüppell, during his extensive travels in East Africa, seems never to have come across it, and does not mention it in any of his publications.

The first author after its describers to recognize its existence was Heuglin, who in 1877, in the second volume of his ‘Reise in Nordost-Afrika,’ writes of this species, the name of which he had previously misapplied to another Gazelle, and gives a coloured figure of its head and figures of two pairs of its horns. Heuglin met with G. leptoceros in the Libyan desert of Egypt, near the Natron Lakes and the Fayoum, where he states its Arabic name is “Abu el haráb.”

Sir Victor Brooke had never seen specimens of this Gazelle, and in his ‘Monograph’ relies mainly upon Heuglin’s description.

So matters remained until recent years, when examples of this Gazelle, or of a very closely allied form, turned up unexpectedly from a new quarter.

Loche, Lataste, and other authorities on the zoology of Algeria had mentioned the existence far in the interior of that country of an Antelope called “El Rim,” and examples of the horns of a problematical Gazelle called “El Reem” had been brought to England from the shops at Biskra. In 1894 an enthusiastic sportsman and naturalist, Sir Edmund Loder, F.Z.S., resolved to make a serious attempt to discover this mysterious animal, and proceeded to Algeria for that purpose. We cannot do better than transcribe for our readers Sir Edmund’s own account of the results of this successful expedition, which was read before the Zoological Society of London on June 5th, 1894:—

On his return home Sir Edmund Loder submitted his series of specimens of the Gazelles obtained during this and his former journeys in Algeria to Thomas, who, at the same meeting of the Zoological Society at which Sir Edmund’s notes were read, proposed to refer his examples of the “Reem” to a new species to be called Gazella loderi, after the energetic traveller who first made known its existence in Algeria.

Mr. Pease, in his notes on the Antelopes of Eastern Algeria published in the Zoological Society’s ‘Proceedings’ for 1896, gives us the following additional information:—

[143]

Mr. Pease also points out that in the “Rhime” the horns in their main outline form a long evenly tapering V, whilst in the Admi the horns are more inclined to be parallel, and towards the points usually take an inward and forward turn, as shown in the diagrams (Fig. 66, a, b) which by the kindness of the Zoological Society we are able to reproduce. The annulations, also, Mr. Pease states, are deeper and more marked in the Admi, and stop more abruptly towards the points than in the “Rhime.”

Fig. 66.

Diagram of horns of Rhime (a) and Admi (b).

(P. Z. S. 1896, p. 814.)

This Gazelle occurs also in Southern Tunis, as we learn from Mr. J. S. Whitaker, F.Z.S., who has written the following notes on it in the Zoological Society’s ‘Proceedings’:—

In the spring of 1895 Sclater was in Egypt, and convinced himself that besides G. dorcas, of which there were many specimens in the Zoological Gardens at Gizeh, there were examples of both sexes of another species belonging to the group of G. leptoceros and G. cuvieri, stated to have been obtained from the Arabs of the Western Desert (cf. P. Z. S. 1895, p. 400). On receipt in London of a skin and skull of this Gazelle subsequently sent to him by Mr. Jennings-Bramley, Sclater, after comparing it with typical specimens of Gazella loderi in the British Museum, pronounced them to be of the same species. Mr. Jennings-Bramley also supplied Sclater with some excellent notes on the mode of capture of this Gazelle by the Arabs of the Western Desert of Egypt, from which we extract the following passages:—

In August 1896 a fine adult living female of this Gazelle was received by the Society as a present from Mr. A. R. Birdwood, of Cairo—no doubt obtained in the same locality as that explored by Mr. Bramley. Mr. Birdwood wrote subsequently to Sclater concerning this Gazelle as follows:—

So far as we know, besides the original specimens of G. leptoceros received at Paris in 1884, the female presented to the Zoological Society by Mr. Birdwood is the only example of this Gazelle that has reached the Menageries of[147] Europe alive. By the kindness of the Zoological Society we are able to give a copy of Mr. Smit’s drawing of the head of this animal.

Fig. 67.

Front view of head of a female Loder’s Gazelle.

(P.Z.S. 1896, p. 781.)

But the identity of the Egyptian Gazella leptoceros with the Algerian G. loderi is perhaps not yet exactly certain, although we have combined the English name of the latter with the scientific name of the former.

On comparing specimens from Tunis and Algeria with others from Egypt, the size of the former is slightly greater, the markings are even less defined than in Egyptian examples, the horns are less closely ringed, the nasal bones are markedly longer, the nasal opening is both longer and broader, and the premaxillæ articulate less broadly with the sides of the nasal bones.

[148]

An old male is 26 inches in height at the withers, and the skull-measurements of the type are:—Basal length 6·75 inches, greatest breadth 3·35, muzzle to orbit 4.

These differences seem to be quite constant, so far as we have materials for comparison, and we therefore think that as the Algerian form has had a name given to it, it may be provisionally retained as a subspecies, at least until these characters are shown to be variable. The accompanying figure, for the use of which we are indebted to the kindness of the Zoological Society, gives a side view of the skull and horns of the Algerian form.

Fig. 68.

Skull of Gazella leptoceros loderi, ♂.

(P.Z.S. 1894, p. 471.)

Our representations of this Gazelle (Plate LXIII.) have been prepared by Mr. Smit—that of the male (front figure) from a mounted specimen in the[149] British Museum obtained by Mr. J. I. S. Whitaker in the Tunisian Sahara, and presented by him in 1894; that of the female from the example from Egypt living in the Zoological Society’s Gardens.

In the British Museum are likewise the typical skin and horns of Gazella loderi obtained by Sir Edmund Loder in the desert about a hundred miles south of Biskra, some frontlets and horns from Biskra, presented by Mr. Rowland Ward, and a skin and skull of a female from Tunis, presented by Mr. J. I. S. Whitaker along with the male now mounted. The example of Gazella leptoceros typica sent to Sclater by Mr. Birdwood is also now in the National Collection. We have also to thank Dr. J. Anderson, F.Z.S., for the loan of a skin and skull of an old male of the Egyptian form of this Gazelle obtained near the Natron Lakes in Egypt.

May, 1898.

[150]

THE BOOK OF ANTELOPES, PL. LXIV.

Wolf del J Smit lith

Hanhart imp.

The Isabella Gazelle.

GAZELLA ISABELLA.

Published by R.H. Porter.

[151]

94. THE ISABELLA GAZELLE.
GAZELLA ISABELLA, Gray.
[PLATE LXIV.]

Height at withers about 25 inches. General colour pale fawn, rather variable in tone, sometimes tending towards brownish. Light lateral band very indistinct; dark band generally fawn like the back, occasionally darker, almost smoky brown. Central dark facial band deep rufous, a darker nasal patch often developing in old individuals. Light facial streak well defined, white. Pygal band almost obsolete, little or not darker than the back.

Skull rather variable in the shapes of the nasal bones and premaxillæ. That of a male measures 6·55 inches in basal length, greatest breadth 3·1, muzzle to orbit 3·65.

Horns of male, as well figured by Blanford under the name of G. dorcas, evenly diverging and curving backwards for four-fifths their length, their tips strongly bent inwards and upwards nearly or quite to a right angle.

[152]

Female. Similar to the male, but the horns slender, scarcely ridged, their tips curved inwards rather than upwards; in length nearly equal to those of the male.

It is possible that the Gazelles described and figured by Lichtenstein in the first part of his ‘Darstellung der Thiere’ as “Antilope dorcas,” which were stated to have been procured by Hemprich and Ehrenberg in Sennaar, may have belonged to the present species. Sundevall certainly considered them to be referable to a species distinct from the true Gazella dorcas, and proposed to call them “isidis,” from Lichtenstein’s vernacular name “Isis Antelope.” But this identification is by no means certain, and, at all events, the name “isabella,” under which this Gazelle was shortly diagnosed by the late Dr. Gray in 1846, will take precedence of Sundevall’s appellation. Gray’s description is very short, and does not allude to the shape of the horns, which are one of the most characteristic features of this species. His type specimen is still in the British Museum. It is an immature male, mounted, and stated to have been received from “Abyssinia,” though Gray in later papers gives “Egypt” and “Cordofan” as the localities for his G. isabella.

Heuglin, in his various memoirs on the Antelopes of N.E. Africa, did not keep G. isabella separate from G. dorcas, and united their localities. Sir Victor Brooke, in his monograph of the Gazelles, though he divides them and says that “amongst the smaller Gazelles no two species could produce two more dissimilar animals than typical specimens of G. dorcas and G. isabella,” states his conviction that “every intermediate degree between them will be found represented in intermediate localities.” It is indeed true that G. isabella is a very inconstant species and requires further careful study.

There can be no doubt that Dr. Blanford’s Gazella dorcas, in his volume on the ‘Geology and Zoology of the Abyssinian Expedition,’ is what we here call G. isabella. The figure of its horns (op. cit. plate i. fig. 1) shows the characteristic twist inwards at the upper end. Moreover, a skull of a male (from Zoulla) and a skull and skin of a female (from Amba), obtained by[153] him during the expedition, and now in the British Museum, are evidently referable here. Mr. W. L. Sclater has also catalogued four heads in the Calcutta Museum, obtained by Mr. Blanford on the same occasion, as G. Isabella. Mr. Blanford gives us the following field-notes on the present species:—

In our efforts to obtain further information about the Isabella Gazelle, we did not fail to apply to the officers of the Anglo-Egyptian garrison at Suakin for a set of specimens of it for the National Collection. In reply to our requests Major Sparkes, Surgeon-Capt. Fleming, and Lieut. Carleton were kind enough to send to the British Museum five examples of it; but we cannot say that the examination of these specimens has enabled us altogether to understand this very difficult species. Of the five examples from that locality, three have and two have not a black nasal patch, while the dark lateral band in some is fairly distinct and in others almost obsolete. It is thus evident that these characters, of systematic importance elsewhere, are not, in G. isabella, even of local constancy.

Among the Gazelles registered in the Zoological Society’s ‘List of Animals’ (1896) as received during the past twelve years there have been several which, doubtless, should have been referred to the present species, but have been entered under Gazella dorcas. Amongst these may be specified an example presented by Commander W. Crofton, R.N., in July 1890 (specimen e), a female presented by Col. Holled Smith, C.B., in July 1892 (specimen g), and a pair (h, i) received on deposit in May 1894.

After the arrival of these specimens from Suakin, Mr. A. Thomson, the Head-Keeper, called Sclater’s attention to their differences from the ordinary G. dorcas. They were of a more reddish colour, and had a broad and somewhat distinct side-stripe and a blackish nasal spot, in addition to other smaller differences.

[154]

Besides the specimens of this Gazelle in the British Museum which we have already referred to, there is a mounted male from the Anseba River, formerly in Sir Victor Brooke’s collection, and presented to the Museum by Sir Douglas Brooke. We believe that our coloured figure (Plate LXIV.), which was prepared by Mr. Smit under Sir Victor’s superintendence, was taken from this specimen.

May, 1898.

Fig. 69. Fig. 69a.

Heads of Isabella Gazelle, ♂ & ♀.

(From specimens in B. M.)

THE BOOK OF ANTELOPES, PL. LXV

Wolf del. J. Smit lith.

Hanhart imp

The Muscat Gazelle.

GAZELLA MUSCATENSIS.

Published by R. H. Porter.

[155]

95. THE MUSCAT GAZELLE.
GAZELLA MUSCATENSIS, Brooke.
[PLATE LXV.]

Size small, height at withers 21–22 inches. General colour dark rufous fawn, darker than in any other species except G. arabica, which it much resembles in colour. Light lateral band scarcely or not perceptible; dark lateral band blackish, its upper edge little defined. Central facial band dark rufous, a distinct blackish patch over the nasals; light facial streak narrow, well defined, dark band below it not defined from the general colour of the cheeks. Knee-tufts present, brownish. Limbs darker in colour than usual, being only white on the inner sides of the forearms and thighs.

Skull very similar to that of G. isabella. Premaxillæ scarcely touching nasals. Basal length in an old female 5·7 inches, greatest breadth 2·9, muzzle to orbit 3·6.

Horns of males curved like those of G. isabella, but decidedly shorter, not or little longer than the skull.

Female. Similar to the male, but the horns slender, scarcely ringed, nearly as long as those of the other sex.

On the 15th of August, 1873, the Zoological Society of London received as a present from Major C. B. Euan Smith (now Col. Sir Charles B. Euan Smith, K.C.B.) a male Gazelle which he had brought with him from[156] Muscat. On September 20th of the same year a female, obviously of the same species and obtained at the same place, was received by the Society on deposit from Mrs. Harris, then of Limefield, Kirkby Lonsdale. Sir Victor Brooke having then lately published his well-known monograph of the Gazelles in the Zoological Society’s ‘Proceedings,’ and being specially interested in the group, Sclater lost no time in calling Sir Victor’s attention to these animals, with which he was much delighted. Sir Victor described them as belonging to a new species at the meeting of the Zoological Society on Feb. 27, 1874, under the name Gazella muscatensis, and pointed out the clear differences which separated them from G. arabica, which up to that time he had believed to be the only Gazelle met with in any part of Arabia.

Fig. 70. Fig. 70 a.

Heads of Muscat Gazelle, ♂ & ♀.

(From specimens in B. M.)

Instead of the massive, nearly straight, non-lyrate horns of G. arabica, Sir Victor showed that the new species had rather slender horns, compressed[157] from side to side and distinctly lyrate, with their points turned boldly forwards and inwards. In general appearance also the Muscat Gazelles differed from G. arabica in their long and soft coats of a silvery-grey colour, instead of the short close-set pelage of a rich grizzled bay. From G. dorcas, to which they bore more resemblance, the Muscat species was recognizable by its smaller size, its different colour, and by the intensity of the facial and lateral markings.

Sir Victor’s paper on this new Gazelle was illustrated in the ‘Proceedings’ by a good coloured plate drawn by Keulemans, in which, however, the general colour is made rather too dark.

In 1874 a second male specimen of this well-marked species was presented to the Zoological Society’s Menagerie by Mr. J. H. Bainbridge, and in October 1881 a pair of the same Gazelle were presented by the late Lord Lilford. These last bred a young one, which was born in the Society’s Gardens on the 6th of March, 1882.

No more examples of the Muscat Gazelle reached the Regent’s Park after this date until 1894, when a female of this species was obtained “in exchange,” and a pair were received “on deposit” from the Hon. Walter Rothschild, F.Z.S. The male of this pair is still living in the Society’s Menagerie.

The only other specimens of the Muscat Gazelle ever received in Europe, so far as we know, are five examples in the British Museum, presented to that Institution by Dr. A. S. G. Jayakar, of whose many and valuable contributions to science we have already spoken[14]. Among the several consignments of the Mammals of Oman sent home by Dr. Jayakar, of which Thomas has given an account in the Zoological Society’s ‘Proceedings’ for 1894, were five examples of this Gazelle collected in 1892 and 1893 in several localities in Oman—Khode and Barkah-al-moze, and in Sharkeeyeh, the eastern part of that country. These specimens agree very closely with the type of the species as described by Sir Victor Brooke, which is also in the National Collection.

Our figures of both sexes of this Gazelle (Plate LXV.) have been prepared by Mr. Smit from the specimens in the British Museum.

May, 1898.

[158]

THE BOOK OF ANTELOPES, PL. LXVI.

Wolf del. J. Smit lith.

Hanhart imp.

Heuglin’s Gazelle.

GAZELLA TILONURA.

Published by R.H.Porter.

[159]

96. HEUGLIN’S GAZELLE.
GAZELLA TILONURA (Heugl.).
[PLATE LXVI.]

Height at withers about 27 inches. General colour deep sandy. Central facial band but little more rufous than the back; no black patch on muzzle. Light facial streak scarcely or not perceptible on sides of muzzle; the area round the eye dull whitish, not sharply defined. Back of ears scarcely lighter than nape. Light lateral band present, not strongly defined. Dark lateral band black, strongly marked, though narrower than in G. thomsoni; a sandy line present between it and the white of the belly. No dark pygal band. Tail sandy at base, the remainder black. Knee-brushes present, dark sandy.

Horns not, or little, longer than the head, lyrate, parallel at base, curving outwards above and then abruptly twisted inwards towards each other at the tip, the ends each forming a sharp hook, similar to that found in G. soemmerringi, but even more strongly bent inwards.

[160]

Fig. 71. Fig. 72.

Heads of Heuglin’s Gazelle, ♂ & ♀.

(From specimens in B. M.)

What little we know of this Gazelle is chiefly due to the researches of the late Baron Theodor von Heuglin, an energetic collector and observer of the Mammals and Birds of North-eastern Africa, whose name we have already had frequent occasion to mention in the pages of this work. In the absence of any better designation, we have selected “Heuglin’s Gazelle” as its English name, which is so far applicable that, besides being its first describer, Heuglin is the only naturalist that has recorded observations on it as met with in its native wilds. Heuglin passed several months in the fertile territory of Bogos, north of Abyssinia (now, we believe, included in the Italian colony of “Eritrea”), when attached to the German expedition sent out in search of the much-lamented traveller Dr. Eduard Vogel. He thoroughly explored this country, which is traversed by the River Anseba,[161] and discovered many new birds and mammals, which were subsequently described in his various works. Amongst the mammals was the present species of Gazelle, which he met with only “on the bushy plains round Ain-Saba from 3000 to 5000 feet above the sea-level, in small families of from three to six individuals.” In his original description Heuglin called this Gazelle Antilope melanura, but subsequently altered its specific name to “tilonura,” there having been already an Antilope melanura of Bechstein, which term is, however, a useless synonym of the Oribi (Ourebia scoparia). We have not been able to discover what the term “tilonura” means, but follow the change, which has been adopted by Sir Victor Brooke and other authors.

Little more, we regret to say, can be told of this beautiful species, which is readily distinguishable amongst its congeners by its broad black lateral stripe and lyre-shaped horns with incurved points. Sir Victor Brooke gave a figure of it from a stuffed specimen in his own collection to illustrate his article on the Gazelles in the Zoological Society’s ‘Proceedings’ for 1873. Our figure (Plate LXVI.), which was prepared by Smit under the direction of Sir Victor Brooke, was probably taken from the same specimen, now in the British Museum, to which it has been presented by Sir Douglas Brooke. It was obtained in Bogosland by Essler about 1872. Four other examples of this Gazelle procured at the same time by the same collector are also in the National Collection. From two of these the accompanying illustrations of the head and horns (figs. 71 & 72, p. 160) have been prepared by Mr. Smit.

May, 1898.

[162]

THE BOOK OF ANTELOPES, PL. LXVII.

Wolf del. J. Smit lith.

Hanhart imp.

The Red-fronted Gazelle.

GAZELLA RUFIFRONS.

Published by R. H Porter

[163]

97. THE RED-FRONTED GAZELLE.
GAZELLA RUFIFRONS, Gray.
[PLATE LXVII.]

Size medium, form comparatively rather stout. General colour deep sandy rufous, brightening into rich rufous on the forehead and muzzle. Nose-spot blackish. White facial streaks well defined, dark cheek-stripes rufous. Light lateral band broad, sandy buff, about the colour of the shoulders, sharply defined from the colour of the back; dark lateral band narrow, black, sharply defined, very prominent, succeeded below by a narrow edging of sandy. Knee-tufts absent. Tail blackish, except just at the base above, where it is sandy.

Skull of medium build; nasals rather broad and short, broadened behind; nasal opening long and narrow, the upper line of the premaxillæ straighter and less concave than usual. Dimensions of a slightly immature male:—Basal length 7 inches, greatest breadth 3, muzzle to orbit 3·95.

Horns rather short in proportion to the size of the animal, evenly[164] divergent, slightly curved backwards below and forwards above, heavily ringed except for the terminal two or three inches.

Female. Similar to the male, but the horns straight, slender, and less ringed; those of an adult rather less than six inches in length.

The existence of a Gazelle of the group allied to Gazella dorcas in West Africa was first made known to us by the authors of the great folio work called ‘Histoire Naturelle des Mammifères,’ issued at Paris, in which the figures were taken mostly from living specimens. In one of the early livraisons of this publication, in which French names only are primarily used, this species was referred to the “Kevel” of Buffon, and under this name a young male was figured, stated to have been brought to France from Senegal. In a later livraison two young specimens of the same species, also from Senegal, were figured as the “Corine” of Buffon, which was declared to be identical with the “Kevel.” We have, however, already shown (under the head of Gazella dorcas) that both these terms of Buffon are referable to the last-named species, and that neither these terms nor the scientific names founded upon them can be properly used for any other species. Hence it follows that the first scientific name that can be employed for this Gazelle is Gazella rufifrons of Gray, under which term it was curtly described by that author in the ‘Annals and Magazine of Natural History’ for 1846. Shortly after that period the same species was figured in ‘Gleanings from the Knowsley Menagerie’ by Waterhouse Hawkins, whose well-drawn plate contains portraits of two males, a female, and a young one of the present species.

As the drawings in the ‘Gleanings’ were not in all cases taken from animals living in Lord Derby’s Menagerie, some of them having been prepared from specimens in the British Museum, it is nearly certain that this was the case in the present instance. It will be observed that Gray in his description mentions almost exactly such a series as being in the Museum as is drawn in the ‘Gleanings,’ and it is not probable that a similar set should also have been living at Knowsley at the same time.

All the specimens mentioned by Gray, with one exception, are still in the National Collection, and, as “co-types” of the species, show clearly to what animal the name “rufifrons” should be applied.

[165]

In 1847, in his excellent essay on the “Pecora,” the late Professor Carl J. Sundevall established an “Antilope lævipes” basing it primarily on the figures of Geoffrey St.-Hilaire and Fr. Cuvier in the ‘Histoire Naturelle des Mammifères,’ of which we have already spoken. It is obvious, therefore, that Sundevall’s term “lævipes” is an absolute synonym of “rufifrons,” though he gives as his principal example a specimen from Sennaar in the Stockholm Museum. But, nevertheless, there would appear to be a closely allied species living in East Africa, which Heuglin, in his first work (‘Antilopen und Büffel’), referred to Antilope leptoceros, but which he subsequently in 1877 (‘Reise in Nordost-Afrika’) treated of under Sundevall’s name “Antilope lævipes.” Heuglin states that this Gazelle, of which the native name is “Abu-el-Harabat,” is found in Nubia, Kordofan, Sennaar, and Taka, as also near Suakin and on the plains of the Beni Amer, ascending to a height of about 1500 feet above the sea-level. Heuglin specially alludes to the want of knee-tufts in his Antilope lævipes, which is the case also in Gazella rufifrons. What this East-African Antelope of Sundevall and Heuglin may be we are quite unable to decide, not having been able to examine East-African specimens. It is possible, however, that it may be Gazella rufina, which we shall presently speak of.

Of the examples of this Gazelle formerly in the Knowsley Menagerie we have already written. The Zoological Society of London have also, on several occasions, received living examples of this species from the West Coast of Africa. The first of these recorded of late years was purchased in August 1865, and the second, a female, in 1869. In June 1895 a female Gazella rufifrons was placed under the Society’s care by the Hon. W. Rothschild, F.Z.S., and is still living in the Gardens. In May of last year a fine living pair of this species were received by the Zoological Society from a London dealer and subsequently purchased. It is from this pair that the accompanying figures of both sexes of this beautiful Gazelle (Plate LXVII.) have been prepared by Mr. Smit. These animals are still living in the Society’s Gardens. They stand about 24 inches in height at the shoulder, the male being slightly the taller of the two. Both male and female have a slight blackish nose-spot, as shown in our figures. The knee-brushes are quite imperceptible in these three specimens, and the knees are perfectly smooth.

May, 1898.

[167]

98. THE RUFOUS GAZELLE.
GAZELLA RUFINA, THOS.

Closely similar in all respects to G. rufifrons, but decidedly larger, the colour throughout particularly rich and brilliant. Light facial streaks not white but sandy, and little different to the general colour of the cheeks. In the single specimen known (a tanned skin), the hairs along the centre of the back and on the sides of the rump are peculiarly waved, but how far this may be due to the preparation of the skin we are unable to say. Knee-brushes dull rufous.

Skull conspicuously larger and heavier than that of G. rufifrons. Anteorbital fossæ larger and deeper. Nasal opening shorter and broader in proportion to the general size, and the upper line of the premaxillæ more curved. Basal length of the type, an old male, (circa) 8·05 inches, greatest breadth 3·86, muzzle to orbit 4·9.

Horns of male short and heavy, their curvature and proportional length about as in G. rufifrons; those of the type are 11·6 inches in length on their anterior curve.

After his return from Algeria in 1894, Sir Edmund Loder, as we have already stated, placed the whole of his series of specimens of Gazelles from[168] that country in Thomas’s hands for examination. Besides examples of the two previously known Algerian species—Gazella dorcas and G. cuvieri—Thomas found that two other species, hitherto apparently unrecognized, were represented in the collection. Of one of these, which Thomas named G. loderi, we have treated under the head of G. leptoceros. Of the other, to which we now propose to refer, only a single specimen, consisting of a skin and a skull, was in the series. This, however, was of so entirely a different character from the three others above-mentioned that Thomas found it necessary to refer it to a new species, which he proposed to call G. rufina, from its generally bright, rich rufous colour. The specimen not having been obtained in the field, but having been purchased at a shop in Algiers (in the spring of 1877), cannot have any certain locality assigned to it; but in all probability it was brought from somewhere in the interior, as it is hardly likely that such an object would have been imported into Algiers from a distant country.

Fig. 73.

Skull of Rufous Gazelle.

(P. Z. S. 1894, p. 468.)

There can be no doubt that, as stated above, Gazella rufina is most nearly allied to G. rufifrons of Senegal, from which, however, we have already stated its points of difference. Sir Edmund Loder has had a water-colour[169] drawing of the typical specimen prepared by Smit, which has much assisted us in our study of it. It is obviously a larger and more richly coloured animal than G. rufifrons, and its skull, of which, by the kindness of the Zoological Society, we are enabled to give the figure that originally appeared in their ‘Proceedings’ (fig. 73, p. 168), is remarkable for its stout and heavy build and short nasal opening. The only conjecture we can make respecting this somewhat problematical species is that it may possibly be the same as Gazella lævipes of Sundevall and Heuglin from North-east Africa, of which we have spoken in our article on Gazella rufifrons. It may be remarked that Sundevall, in his account of Antilope lævipes (K. Vet.-Akad. Handl. 1845), has noted the existence, in the Paris Museum, of specimens from Algeria of a Gazelle which he considers identical with his species, form α (from Sennaar), and which he separates from form β (from Senegal) on account of the want of the blackish nose-spot. But even if such shall turn out eventually to be the case, the present species will still retain the name “rufina,”—“lævipes,” as we have already pointed out, being correctly treated of as a synonym of G. rufifrons.

Sir Edmund Loder has most liberally presented the typical specimen of Gazella rufina to the British Museum.

May, 1898.

[170]

THE BOOK OF ANTELOPES, PL. LXVIII

J. Smit lith.

Hanhart imp.

Thomson’s Gazelle.

GAZELLA THOMSONI.

Published by R. H. Porter.

[171]

99. THOMSON’S GAZELLE.
GAZELLA THOMSONI, Günth.
[PLATE LXVIII.]

Height at shoulders about 25 inches. General colour deep sandy rufous, all the markings well developed and sharply defined. Central facial band deeper rufous, a black patch present on the top of the muzzle. Light facial streaks pure white, extending quite to the muzzle and surrounding the eye; darker cheek-streaks blackish. Light lateral band present, though not very much paler than the back. Dark lateral band deep black, greatly developed, very broad, sharply defined above and below, extending on to the outer side of the elbow; no sandy streak between it and the white of the belly. Pygal band small, blackish. Ears rather short. Tail sandy at its base above, the remainder black. Knee-brushes present, sandy or whitish.

Skull with short, broad, and quadrangular nasals, nearly as broad as long; premaxillæ straight, scarcely concave above, broadly articulating with nasals. Basal length of skull of a male 7·7 inches, greatest breadth 3·6, muzzle to orbit 4·25.

[172]

Horns rather like those of G. bennetti on a large scale, little divergent, sometimes even quite parallel, evenly but very slightly curved backwards for seven-eighths of their length, their tips gently recurved upwards and forwards. In length, over the front curve, they attain to about 14 or 15 inches, the record being 15½.

Female. Similar to the male, but with the horns rudimentary, much smaller than in other African Gazelles; only from 3 to 6 inches in length, of about the thickness of a cedar pencil, smooth and unridged, and in direction crooked and irregular.

Fig. 74.

Horns of Thomson’s Gazelle, ♂.

(Ann. Mag. N. H. (5) xiv. p. 427.)

The name of Joseph Thomson, the African traveller, will always rank among the foremost of those who in the second half of this century have striven to open the Dark Continent to civilized man, and have lost their lives at an early age by violence or disease in consequence. Thomson was, moreover, one of the very few amongst African explorers who had never shed the[173] blood of a native, nor even, we believe, during his many journeys, fired a shot in self-defence. Thomson’s name has been worthily associated with the present Gazelle, of which he was the discoverer during his expedition through Masai-land to Lake Victoria in 1883 and 1884, and of which he first brought back specimens to Europe.

Thomson, as we are informed by Dr. Günther, who described this Gazelle as Gazella thomsoni in June 1884 from two frontlets presented by Thomson to the British Museum, met with it on his way up the country from the plains near Kilimanjaro to Lake Baringo, at various elevations under 6000 feet. Dr. Günther, we may remark, in his description and figure of these horns fell into a not unnatural error in treating the more slender pair (fig. 74a, p. 172) as those of a female. But, as we have already stated, the horns are always abnormally small in the doe of this Gazelle, and sometimes, it is said, altogether wanting. The slenderer pair of horns shown in Dr. Günther’s figures, which we have been kindly allowed to reproduce in this work, are, like the stouter pair, doubtless those of a male.

In his volume ‘Through Masai-land,’ Thomson does not appear to have made any reference to this Gazelle, except by repeating the figures of the horns (p. 536) already published by Dr. Günther. Thomson had intended, we believe, to put his notes on the animals and plants collected and observed during this expedition into an Appendix, which, however, from pressure of other matters, was never written.

After Thomson himself, the next earliest information obtained concerning this Gazelle appears to be that collected by Sir John Willoughby’s hunting-party in 1886–87. In the Appendix to ‘East-Africa and its Big Game,’ Mr. Hunter writes of it as follows:—

In the first volume of ‘Big Game Shooting,’ of the Badminton Library, Mr. F. J. Jackson, than whom no one can be better qualified to speak of[174] East-African Antelopes, gives us his experiences with the present species in the following words:—

Fig. 75.

Front view of head of Thomson’s Gazelle, ♀.

(Neumann’s ‘Elephant-Hunting,’ p. 11.)

Writing subsequently in the Zoological Society’s ‘Proceedings’ on the Antelopes of the Mau District of British East Africa, Mr. Jackson says that this Gazelle does not, as he believes, extend beyond a few miles north of Lake Nakuru. He adds that “the females are horned, whatever may be said to the contrary.” With this view Mr. Arthur H. Neumann, one of our most recent explorers in British East Africa, quite agrees. In his lately published volume on ‘Elephant-Hunting’ in that Protectorate, Mr. Neumann gives a figure of the head of a female Thomson’s Gazelle, which, by his kindness and that of his publishers, Messrs. Rowland Ward & Co., we are enabled to reproduce (fig. 75, p. 174): this shows that the horns are present in that sex of G. thomsoni, though in a much more dwarfed condition than in most of its allies. Mr. Neumann also claims to have met with this Gazelle much further north than its range is usually held to extend. This was in the district of Kisima, south of the Lorogi Mountains and north of Lake Naivasha.

Col. Lugard, who has had great experience in East-African game-shooting, has stated[15] that he had never met with a horned female of Thomson’s Gazelle. A letter lately received from our much-valued correspondent, Mr. S. L. Hinde of the B.E.A. Medical Service, in response to enquiries on this point, endeavours to explain this diversity of opinion as follows:—

Passing on to German East Africa we find, from Dr. Matschie’s volume on the Mammals of that country, that Thomson’s Gazelle was met with by Herr Oscar Neumann at various points on his journey north from Tanga to the Victoria Lake. Mount Gurui, Lake Manyara, Ndalalani on the Natron Lake, and the district between Guasso Nyiro and Ngare Dobasch are specially mentioned, but north and west of the latter locality Herr Neumann did not find it. It was also obtained near Mount Meru by Herr v. Höhnel, and in Northern Irambi by Herr Stuhlmann.

We are not aware that examples of Thomson’s Gazelle have ever been brought to Europe alive; but it would appear that young individuals of this species are often captured and kept in confinement in British East Africa. The late Capt. B. L. Sclater, R.E., who made the road from Kibwesi to Port Victoria in 1895–97, in his letters written home refers to several such cases, and Lieut. G. E. Smith, R.E., who was second in command of Capt. Sclater’s party, kindly sends us the following notes about them:—“Early in January 1896 Capt. Sclater obtained from the Masai two young Thomson’s Gazelles, which were fed by hand and throve well. They became quite tame and used to run about the camp and play with a puppy. One of them subsequently ran away, but the other remained with the camping-party for nearly a year.” At Kikuyu, Lieut. G. E. Smith saw a pair of Thomson’s Gazelles then fully grown, having been in possession of Mr. Hall, the Resident, for more than two years. The buck was rather fierce and apt to attack natives with his horns. Major Eric Smith, we are told, also generally[177] had two or three of these Gazelles scampering about his station at Naivasha very tame.

Our figure of the male of this Antelope (Plate LXVIII.) has been prepared by Mr. Smit from a mounted specimen in the British Museum presented by Mr. F. J. Jackson. There are also in the Museum a second mounted specimen of a male presented by Mr. H. C. V. Hunter and the skull of a male from Lake Naivasha presented by Col. Lugard. Besides these the Museum contains two frontlets with horns, being the typical specimens obtained by Thomson, and some skins and skulls of both sexes from Machakos, presented by Mr. S. L. Hinde, as already mentioned in his letter.

May, 1898.

[178]

THE BOOK OF ANTELOPES, PL LXIX

Wolf del J Smit lith

Hanhart imp

Grant’s Gazelle.

GAZELLA GRANTI.

Published by R. H. Porter

[179]

100. GRANT’S GAZELLE.
GAZELLA GRANTI, Brooke.
[PLATE LXIX.]

Size large; height at withers about 34 inches. Fur close and short. General colour fawn, rather variable in tone. Lateral bands, both light and dark, usually very indistinct, often scarcely perceptible; but, on the other hand, occasionally well developed, especially in young animals. Central facial band richer fawn, approaching rufous; a brownish spot present on the muzzle. Light facial streaks white, sharply defined, running up over the eyes to the horns. Below these the dark facial bands are almost imperceptible, scarcely or not at all darker than the fawn-coloured cheeks and neck. Rump with the white of the hams very broad, extended upwards,[180] and uniting across the base of the tail, so that the latter is quite separated from the dark body-colour; laterally the white penetrates angularly into the body-colour, overhanging the top of the pygal band, which is generally well-defined. Outer sides of limbs fawn, without blacker markings. Knee-brushes present, dark fawn. Tail above white for its basal half; black and crested terminally.

Skull stout and heavy, nasal opening broad. In that of an adult male the basal length is 9·75 inches, greatest breadth 4·4, muzzle to orbit 5·3.

Horns very long, longer and more powerful than in any other Gazelle; evenly but slightly curved backwards below, and gently recurved forwards terminally; sometimes but slightly divergent, but more often, especially in specimens from Kilimanjaro, they spread widely above, approaching each other again at their tips. Their section at base is a long oval, very different from the nearly circular section found in G. soemmerringi.

Female. Similar to the male, but the horns slender, nearly circular in section, more strongly ridged than in the females of most Gazelles; about two-thirds in length of those of the male.

Grant’s Gazelle, which has been appropriately named after one of its discoverers, is pre-eminent, even in this ornamental genus, for its size and elegance, and is, in fact, generally allowed to be one of the most beautiful species of the whole group of Antelopes. Speke and Grant left Zanzibar on the well-known expedition during which the efflux of the Nile from Lake Victoria was discovered, in September 1860. Starting from Bagamoyo on the opposite coast, and passing through Usagara, they arrived about two months later in Ugogo, then under the rule of a native chief called Magomba. It was in December 1860 during their stay at this place, where they were long detained by the drunken chief and his wazir, that Speke first met with the present Antelope. In the ‘Journal’ of his travels Speke tells us that while kept waiting to arrange the amount of his “hongo” he took the time out in the jungles very profitably, “killing a fine buck and doe Antelope of an unknown species.” “These animals,” he continues, “are of much about the same size and shape as the common Indian Antelope, and like them roam about in large herds, the most marked difference between the two being in[181] the shape of their horns, and in their colour, in which in both sexes the Ugogo Antelopes rather resemble the Gazella picticaudata of Tibet, except that the former have dark markings on the face.”

Fig. 76.

Grant’s Gazelle, Ugogo.

(From Speke’s ‘Journal of Discovery,’ p. 61.)

The original woodcut of Speke’s ‘Journal,’ drawn by Wolf, as shown by his initials, we have now the pleasure of reproducing by the kind favour of the publishers of that work. It was no doubt executed under Speke’s direction, and, although not quite satisfactory according to our present knowledge of the animal, gives several views of the shape and size of the horns of Grant’s Gazelle.

Although Speke in his ‘Journal’ writes of the Ugogo Antelope as being undoubtedly new, he was at first evidently by no means certain about this point, as in a letter addressed to Sclater from Kazeh, in February 1861, he referred it doubtfully to Gazella soemmerringi. But in a footnote to this[182] letter (which was published in the ‘Proceedings’ of the Zoological Society for January 1863) Sclater stated his conviction that, so far as could be determined from the rough sketch of the horns which accompanied the letter, the Ugogo Antelope was certainly new, and he added that the late Dr. Gray was of the same opinion.

Fig. 77. Fig. 78.

Heads of Grant’s Gazelle, ♂ & ♀.

(From Mr. Jackson’s specimens.)

The box containing the specimens of the new Gazelle from Ugogo having[183] been unfortunately lost on its journey to the coast, Sclater was unable to give any account of this animal in his paper on the Mammals collected and observed during the East-African Expedition, in the ‘Proceedings’ of the Zoological Society for 1864. The subject therefore was delayed until 1872, when Sir Victor Brooke, who was at that time busy upon the Antelopes, obtained from Col. Grant and Capt. Speke the careful sketches of the heads and skins of this animal which the travellers had made in their note-books. The examination of these sketches confirmed Brooke in the opinion that the Antelope represented by them was undoubtedly new to science; and consequently on April 16th of that year Sir Victor read a communication to the Zoological Society in which he proposed to name the new Gazelle after Grant, Speke’s name in this group of animals having been already commemorated by the Gazella spekei of Blyth and the Tragelaphus spekei of Sclater. Sir Victor gave as good a description of the new Antelope as he could from the notes before him, especially alluding to the extraordinary development of its horns, which attained dimensions nearly double those of any other Gazelle known to him. Brooke’s paper, which is accompanied by a beautiful coloured figure of the new Antelope, prepared by Wolf from Speke’s sketches, states that Col. Grant, who had supplied him with copious extracts from his note-books, informed him that this species was only met with during their expedition in Western Kinyenye, in Ugogo. The country inhabited by it he described as low-lying sandy plains, dotted over in some places with euphorbias, dwarf acacias, and stunted baobabs. The chief peculiarity of this district, owing doubtless to its comparatively low level, was the great accumulation of salt, which had of course a marked effect on the vegetation. Water at all times of the year was very scarce there, and often entirely absent, the little found being brackish and undrinkable.

In 1875 the Zoological Society received as a present from Sir John Kirk, then British Consul-General at Zanzibar, a living female Grant’s Gazelle, which, however, was unfortunately in very poor condition and died shortly after its arrival in London. The acquisition of this animal was announced by Sclater in a report on the additions to the Society’s Menagerie read on November 2nd of that year, and was accompanied by a figure of it drawn from the stuffed specimen by Mr. Smit. So far as we know, this is the only individual of Grant’s Gazelle that has ever reached Europe alive.

It will be observed that the original place of discovery of Grant’s Gazelle[184] was in what is now German East Africa. In this country, according to Herr Matschie, it has an extensive range over the whole western portion of the colony, extending up to Mpapwa, in which district several sportsmen have met with it, and through Irangi on to Lake Victoria on the borders of British territory. Over this country it has been found in many localities by Fischer, Neumann, and Stuhlmann. Throughout these districts it appears to resort principally, as stated by Grant, to the more open sandy plains, which are thinly covered with euphorbias and other trees, especially where a saline vegetation is prevalent.

In the eastern districts of British East Africa, Grant’s Gazelle appears to be not less prevalent in similar spots. Mr. Hunter, in his Appendix to Willoughby’s ‘East Africa,’ tells us that during his expedition in 1888 it was found to be common everywhere in the open plains near Kilimanjaro, one male being generally accompanied by from 10 to 15 females. It was at that time most common in the direction of Useri. The longest pair of horns obtained by the party measured 30 inches in length, but 27 inches were considered to constitute a big head. The best female’s head obtained measured 17 inches along the inside curve. Dr. Abbott, during his expedition to Kilimanjaro, as recorded by Mr. True, obtained a good series of specimens of this Antelope, which are now in the United States National Museum. Capt. F. D. Lugard, in his volumes on ‘The Rise of our East-African Empire,’ writes of Grant’s Gazelle as being “a model of symmetry,” while “its large and massive horns are most beautiful in their curve and are carried most gracefully.” He continues as follows:—

Another excellent authority on the Antelopes of British East Africa, Mr. F. J. Jackson, in a recent article on the subject, writes as follows:—

Dr. Donaldson Smith has recorded the existence of this Gazelle “in great numbers” all along his route homewards from Lake Rudolph on the northern bank of the Tana. But these animals may possibly have belonged to the next species Gazella petersi. Dr. Donaldson Smith also met with Grant’s Gazelle in the Boran country “a little west of 39° W.L.,” and found it “extending as far north as 6° N.L.” Mr. Cavendish likewise obtained examples of this Gazelle during his recent expedition to Rudolphia.

There is a good mounted specimen of an adult male of this Antelope in the Gallery of the British Museum, obtained by Mr. F. J. Jackson in the Kilimanjaro district of British East Africa and presented by him. Mr. Jackson has also given the Museum a fine skull of an adult male from the same locality, and has placed at the service of the Museum three other mounted heads of this Antelope. The National Collection also contains skulls of an adult pair of Grant’s Gazelle from Kilimanjaro presented by Mr. W. Carlisle Fraser, and several skulls and skins of different ages from[186] Machakos, collected and presented by Mr. S. L. Hinde. Besides these some skulls and skins obtained during the Cavendish expedition to Lake Rudolph, and presented by Mr. H. S. H. Cavendish and Mr. H. Andrew, have lately been added to the series.

Our illustration of this beautiful Antelope (Plate LXIX.), which shows the horns of the male in various attitudes, was prepared by Mr. Wolf under Sir Victor Brooke’s directions and drawn upon the stone by Mr. Smit. We are not sure as to the specimens from which it was taken.

September, 1898.

[187]

101. PETERS’S GAZELLE.
GAZELLA PETERSI, Günth.

Size large; height at withers about 33 inches. General colour and markings practically identical with those of the last species, Gazella granti, with the exception that the white rump-patch is of much less extent. This patch, which in G. granti is very broad, and projects prominently forward on each side, overhanging the dark pygal band, and passes across above the tail, separating that organ entirely from the dark dorsal colour, is in G. petersi divided above into two portions by an extension of the body-colour which runs down on to and along the top of the tail. Laterally the white is much narrower, and encroaches much less on the body-colour above, scarcely or not at all overhanging the pygal band.

Skull rather smaller than in G. granti, and the nasal opening rather narrower. Basal length in a male 9·3 inches, greatest breadth 4·2, muzzle to orbit 5·4.

Horns similar to those of G. granti, but smaller and more uniformly parallel, never widely divergent above. Horns of female about two-thirds the length of those of the male, slender and comparatively straight.

In a communication made to the Royal Academy of Sciences of Berlin in October 1879 the former Director of the Berlin Museum, the late Dr. Wilhelm Peters, gave an account of the specimens of Mammals collected in East Africa in 1878 by the well-known traveller Dr. G. A. Fischer. Amongst these were the skull and skin of a young male Gazelle obtained at Gelidja, near the mouth of the Osi and Tana Rivers, on June 27th, and stated to be called there by the Swahilis and Wapakomos “Sala.” Peters added to his paper on this subject an excellent lithographic plate of the skull and horns in question, which he referred, without much doubt, to Gazella granti, remarking, however, that the nose-spot was not well defined, and that the horns were straight at the base and not curved.

Fig. 79.

Skull and horns of Peters’s Gazelle, ♂.

(From one of Mr. Jackson’s specimens in the British Museum.)

[189]

In 1884 Dr. Günther, commenting on Peters’s figure of this supposed Gazella granti, stated that in his opinion it “clearly belonged to a distinct species. It resembled somewhat Gazella thomsoni in the slight degree in which the horns diverge from each other; but their annulated portion was almost straight, and the annuli themselves were much further apart, much fewer in number (about twelve), and lower towards the hinder part of the horn. The base of the bony core showed a bossy swelling, which was different from that of Gazella granti, and entirely absent in Gazella thomsoni.” This species Dr. Günther proposed to call Gazella petersi.

There has, however, been much doubt raised by subsequent authorities as to the validity of this species, and although we have given a separate heading to Gazella petersi we have had considerable hesitation as to whether it ought not to be regarded rather as a local subspecies of G. granti, to which it is closely allied in all essential characters. But since we have as yet seen no specimens absolutely intermediate between the two, so far as the rump-markings are concerned, we have thought it advisable to keep them provisionally separate. At the same time we shall be in no way surprised if such specimens should occur, in which case the two forms will have to be united. As regards the skulls, no constant or tangible difference can be made out, although those of G. petersi can usually be distinguished by their smaller size, narrower nasal opening, and shorter and less divergent horns.

Mr. F. G. Jackson, in ‘Big Game Shooting,’ gives us the following account of this Gazelle and his experiences with it:—

“Gazella petersi (known to the Swahilis also as ‘Sala’) may be a local variety of G. granti rather than a distinct species. It used to be plentiful at Merereni on the British East African coast, and is still found further inland in the Galla country. It is certainly a smaller beast than Gazella granti from Kilimanjaro and Machakos, but in other respects is almost identical, excepting in the shape and size of the horns, which I have never known to exceed 22 inches in length measured along the curve. The horns are also straighter, and have not nearly such a pronounced backward curve as those of G. granti, neither do they diverge towards the points so much, being rarely more than seven or eight inches apart at the widest parts. Gazella petersi is found in the small open plains and open scrub.”

In the gallery of the British Museum there is a good mounted specimen of the adult male of this species, also another younger specimen from the[190] “Mainland opposite Zanzibar,” and a third from “Mombasa,” besides two skulls (♂ and ♀) from “South Somaliland,” all presented by Sir John Kirk. These were formerly labelled G. granti, but must be referred to G. petersi if the species are kept separate. There is also in the Museum a fine skin and skull of a male G. petersi from near Mount Pika-pika, about 40 miles from Mombasa, presented by Mr. L. E. Caine. It would appear, therefore, that the range of G. petersi until lately extended all along the coast of British East Africa, though it has now perhaps been nearly shot out in the southern parts.

The British Museum has likewise three skulls of this Gazelle from the Tana River presented by Mr. F. J. Jackson, to whom we are also much indebted for the loan of two other representative heads of this species from Merereni, south of Formosa Bay.

September, 1898.

[191]

102. THE BANDED GAZELLE.
GAZELLA NOTATA, THOS.

Only known from a single flat skin, without the head. Fur unusually long and shaggy. Size about as in G. granti, and general body-markings as in that species, but all much intensified. Dark and light lateral bands much longer and broader, the former nearly black and reaching forwards on to the shoulder, and backwards nearly to the white rump-mark; the latter pale buff, and succeeded above by a second dark band, lighter than the main lateral band, but distinctly darker than the centre of the back. This second dark band united with the other behind the posterior end of the light band. Pygal band black and very strongly defined.

Horns said by the discoverer to have been like those of G. granti.

In Mr. Arthur H. Neumann’s recently published volume on ‘Elephant-Hunting in East Equatorial Africa’ there will be found an account of his adventurous journey to Lake Rudolph, during which, on more than one occasion, he made his camp for some months at El Bogoi, a place situated east of the Loroghi Mountains, in rather higher than 1° N. lat., which was a favourite station for elephants. While at this place in October 1895 Mr. Neumann, accompanied by his native attendants, made an excursion over the Loroghi range, and encamped close to the edge of the open country on their western slope, at an elevation reckoned to be about 5500 feet above the sea-level, at a place called in the map attached to his narrative Kisima. Here, on taking a stroll into the open, he “shot a brace” of what he at first[192] supposed to be Grant’s Gazelles; but on examination he found that they differed from the Grant’s Gazelles of the other side of the range, and of everywhere else that he had been, “in having longer hair, and dark bands on the sides,” while the “shade on the back” was also deeper than in the common kind. On one of the skins thus obtained and presented to the British Museum (see fig. 80), Thomas has based his subspecies “Gazella grantii notata.”

[193]

For much the same reasons as in the case of G. petersi, that is because no intermediate examples were yet known, we decided, when drawing up our Synopsis of Gazelles (above, p. 69), to give a separate heading to this form.

Fig. 80.

Skin of the Banded Gazelle.

But since our Synopsis was prepared, Lord Delamere has sent home some skins which are, to a certain extent, intermediate between G. granti and G. notata in the development of the dark markings, and which, therefore, support Thomas’s original view that the latter should be looked upon as a subspecies of the former. Yet, on the whole, Lord Delamere’s skins are more like G. granti than G. notata, especially in the character of their fur, so that they may be provisionally assigned to the older known form, leaving the exact status of G. notata to be settled when further materials are available.

Thomas stated, in his original description of G. notata, that this Gazelle had also been obtained by Mr. H. S. H. Cavendish, that traveller having supposed that he recognized in the typical skin of G. notata a Gazelle which he had met with during his journey to Lake Rudolph. This supposition, however, has been proved to be mistaken, as the Gazelle in question when brought home turned out to belong to G. thomsoni—an animal up to that date unknown to Mr. Cavendish. Mr. Neumann, the donor of the typical skin of G. notata to the British Museum, remains, therefore, the only sportsman who has yet met with this handsome form, concerning which we hope to receive further information before long.

September, 1898.

[194]

THE BOOK OF ANTELOPES, PL. LXX.

Wolf del. J. Smit lith.

Hanhart imp.

Soemmerring’s Gazelle.

GAZELLA SOEMMERRINGI.

Published by R. H. Porter.

[195]

103. SOEMMERRING’S GAZELLE.
GAZELLA SOEMMERRINGI (Cretzschm.).
[PLATE LXX.]

Size large, height at withers 35½ inches in an old male of the Somali subspecies. General colour very pale fawn, and very uniform everywhere, as there are neither light nor dark lateral bands nor any pygal bands. Central facial band black or blackish fulvous, contrasting markedly with the white lateral facial streaks. Dark facial streaks also black, but very narrow. Sides of muzzle black, continuous with the central facial band. Back of ears whitish, margined and tipped with black. White of rump very broad and extended, projecting far into the body-colour, which it broadly shuts off from the tail. The latter is white basally, black tufted terminally. Knee-brushes present, whitish or fawn.

Basal length of skull 8·85 inches, greatest breadth 4·2, muzzle to orbit 5·4.

Horns long, nearly circular in section, heavily ringed. In the typical subspecies they are but little divergent for their basal half, but then curve widely outwards above, their tips being again abruptly hooked inwards so as to point almost directly towards each other. In the Somali subspecies their length is greater and their divergence is quite even, not increasing above, so that their upper portions are not nearly so widely separated; their tips also hook rather forwards, and not so directly inwards.

Female similar to the male, but the horns much thinner and less rough, though almost as long as those of the male. In both of the subspecies their curvature is closely similar to that found in their respective males.

This fine Gazelle was one of the many discoveries made in North-east Africa by the great traveller and naturalist Edouard Rüppell, and was first[197] described and figured by Cretzschmar in the ‘Atlas’ which illustrated the zoological portion of Rüppell’s ‘Reise.’ The species was dedicated to Ritter Samuel Thomas von Soemmerring, on the occasion of that savant attaining the fiftieth year of his Doctorate—an event which was celebrated by the Senckenbergian Naturalists’ Society of Frankfort-on-the-Main on the 7th of April, 1828. The original specimens were procured by Rüppell on the eastern slope of the Abyssinian coast-range, and were deposited along with all his other collections in the Senckenbergian Museum. In a résumé of his knowledge of the East-African Antelopes contained in a subsequent work on the Vertebrates of Abyssinia, Rüppell tells us that he only met with this species in the bush-clad valleys of the Abyssinian coast, but that it was said to occur also on the large island of Dahalak off Massowah. It was generally observed in small families, which, however, sometimes congregated into larger herds.

Fig. 81.

Skull and horns of Gazella soemmerringi typica (male).

(From a specimen in the British Museum.)

Heuglin, another good authority on the animals of North-east Africa, informs us that this Antelope extends along the coast of the Red Sea from 20° N. latitude southwards to the Danakil country, and that it is also found in valleys of the Baraka and Atbara in the interior, and extends up to the neighbourhood of Berber, but is not so plentiful here as in the coast-districts.

[198]

Dr. W. T. Blanford, who accompanied the British Abyssinian Expedition of 1867–68 as naturalist, found this Antelope abundant on the coast of the Red Sea near Annesley Bay and Massowah, but states that it never ascends the hills. It was met with principally in the low bush and acacia-scrub in herds varying in size from a few individuals up to one hundred or more. A very large number, he adds, were shot by the sportsmen of the Expedition, who found the flesh excellent eating.

Fig. 82a. Fig. 82b.

Horns of Gazella soemmerringi berberana, ♂ & ♀.

(From specimens in the British Museum.)

Examples of Gazella soemmerringi are recorded by Sundevall as being contained in the extensive collections received by the Stockholm Museum which were made by the Swedish traveller Hedenborg in Sennaar, but the exact localities are not specified.

[199]

So much is all we have to say about the smaller and typical form of this species originally discovered by Rüppell. But about the larger subspecies of Northern Somaliland, which has been named berberana by Herr Matschie, we are able to give greater details from information supplied by the many naturalists and sportsmen who have of late years visited the country which it inhabits. Although it does not appear that there is any discontinuity between the ranges of these two forms, and it is quite likely that intermediate specimens may occur where the two forms meet, we cannot but allow that Herr Matschie was justified in assigning a subspecific name to the Southern form. Not only is the latter a larger and finer animal with longer horns, but the curvature of its horns is, as we have already pointed out, so different that, so far as our experience goes, there can be no difficulty in distinguishing the two forms by this character alone.

Capt H. G. C. Swayne, R.E., in his ‘Seventeen Expeditions to Somaliland,’ writes thus of Gazella soemmerringi:—

Captain R. H. Light, of the Indian Staff Corps, who visited Somaliland in 1891, and has kindly furnished us with some field-notes on its Antelopes,[200] writes that this Gazelle is found there in couples, and also in herds of fifty or more, and generally frequents plains with slight coverts. They are not usually difficult of approach, and out of a large herd he has shot four, one after the other, within a mile of ground, the herd moving off at every shot, but allowing him to approach again. At 150 yards distance or so Capt. Light says it is very hard to distinguish a buck from a doe of this Antelope, the size of the body being the same, and the slight differences in the thickness of the horns and neck being hardly perceptible. Their gait is longer and slower than that of Pelzeln’s Gazelle, but still not awkward like that of the “Gerenuk” (Lithocranius walleri).

Mr. F. Gillett, F.Z.S., who accompanied Dr. Donaldson Smith during the first portion of his journey to Lake Rudolph, and made other expeditions in Somaliland, has favoured us with the following notes on this species:—

To Mr. Alfred E. Pease, M.P., F.Z.S., we are greatly indebted for the following notes on his experiences with this Antelope.—

Of the typical form of this Antelope the British Museum possesses an adult male specimen (mounted, but not exhibited), obtained by Esler in the Bogos country, and purchased in 1873, also a female from Sennaar received from the Stockholm Museum, in exchange, in 1846—probably one of those collected by Hedenborg. It has likewise a young male from Abyssinia procured by Rüppell, the original discoverer of the species, and received in exchange from the Frankfort Museum; a skull of a male obtained by Mr. W. Jesse[203] during the Abyssinian Expedition; and several heads and skins from Suakin collected and presented by Major W. Sparkes, of the Welsh Regiment, now attached to the Egyptian army.

Of the southern subspecies, Gazella soemmerringi berberana, the National Collection possesses a stuffed adult male specimen obtained on the Shebeyli River, Somaliland, and presented by Col. Arthur Paget, who has also given to the Museum two heads (male and female) from the Bourdap Mountains in the same country. In the British Museum there are likewise a mounted head and skins of both sexes of this Antelope collected by Capt. Swayne in Somaliland and presented by Sclater.

The first living examples of Soemmerring’s Gazelle received by the Zoological Society were purchased of a dealer in August 1867. The announcement of their arrival, which appeared in the ‘Proceedings’ for November 14th of that year, was accompanied by a beautiful lithographic plate of the young pair drawn by Mr. Wolf. In 1871 a fine young male example of this Antelope was presented by Mr. Charles McIver, Jun., with the information that the specimen had been obtained in the desert on the Red Sea coast about 100 miles south of Suez. If this was correct, the range of this Antelope would appear to extend considerably further north than we know of upon any other authority. Other specimens of this Antelope were obtained in 1877, 1879, 1892, and 1895, including a fine pair from Suakin presented by Col. Holled Smith, C.B., in July 1892. At the present moment, also, there is a fine male of this Antelope in the Society’s Gardens, obtained by purchase in 1897.

Our representation of this Antelope (Plate LXX.) was drawn by Mr. Wolf under Sir Victor Brooke’s directions, and put upon the stone by Smit. It is that of an adult male example, but we unfortunately do not know from what specimen it was taken.

September, 1898.

[204]

THE BOOK OF ANTELOPES, PL. LXXI

J. Smit del. et lith.

Hanhart imp.

The Red-necked Gazelle.

GAZELLA RUFICOLLIS.

Published by R. H. Porter.

[205]

104. THE RED-NECKED GAZELLE.
Gazella Ruficollis (Ham. Smith).
[Plate LXXI.]

[206]

Size large, form graceful. Colour nearly all over white, the only parts which are of the usual rufous shade being the neck and anterior back, the sides and rump gradually shading off into white. Head and ears white, a faint rufous tinge on the centre line of the face. Neck deep rufous all round, except for a white spot on its anterior surface. Lower part of fore legs faintly rufous, rest of limbs and whole of tail white. Knee-brushes present.

Skull, basal length in an immature male 8·85 inches, greatest breadth 4·15, muzzle to orbit 5·3.

Horns of male thick, curved strongly backwards below, then hooked inwards and forwards above. Horns of female almost equally long, but comparatively thin, slightly curved, and not hooked terminally.

In concluding our account of the true Gazelles we come, at the end of the list, to a small but attractive group of three species which are closely related to one another, and take each other’s places as representatives in the different countries where they have been found. These three Antelopes are certainly nearly allied, and it is by no means improbable that, although all the accessible specimens known to us are distinguishable, intermediate forms will ultimately be found to link them together. It is especially likely that this will prove to be the case with the two western species, G. dama and G. mhorr. All the three species of this group are exceedingly rare in collections, and we have been able to obtain but very little information about them, and very little material for comparison, the British Museum being badly off for specimens of all of them. It is to be hoped, however, that the prospective opening of the Soudan, by France on one side and England on the other, will lead to an increase of our knowledge of this group of Antelopes, and of the many other interesting forms of the great North-African desert.

We will commence our account of these three Gazelles with the one which inhabits the eastern part of the Soudan, where our own countrymen may soon be expected to meet with it.

The first notice of the existence of a species of this form in North-eastern Africa appears to have been given by Lichtenstein, who read a paper on the Antelopes of Northern Africa before the Academy of Sciences of Berlin on March 11th, 1824. Amongst the four Antelopes discussed in this learned[207] treatise, which was mainly based upon the specimens sent to the Royal Collections by the well-known travellers Hemprich and Ehrenberg from Dongola and Sennaar, were several representatives of the present species which Lichtenstein not unnaturally referred to the Antilope dama of Pallas. The same course was pursued by Hemprich and Ehrenberg themselves, who shortly afterwards published full descriptions and figures of it in their ‘Symbolæ Physicæ.’ They inform us that they met with specimens of this Antelope in Southern Dongola in the month of July 1822, and hunted it along with the Addax and Leucoryx, which occurred in the same district. They found it plentiful in herds and easy of access, even without the use of horses. Like the other species mentioned, it feeds principally on the acacias. They did not meet with this Antelope until they arrived at 20° N. lat. going south, after which they found it abundant. The Arabs, who much esteem the flesh and sell it when dried, call it “Addra.” It did not appear to approach the banks of the Nile, but kept entirely to the desert and to the valleys which traverse it, especially to the Chor-el-Lebben.

Not far from the same date another distinguished German traveller and naturalist, Rüppell, whose name we have already had frequent occasion to mention, also met with this Antelope. Rüppell sent his specimens to Frankfort-on-the-Main, where they were described and figured by Cretzschmar on the part of the Senckenbergian Society of Naturalists in 1826. Cretzschmar also referred these specimens to Antilope dama; but Hamilton Smith, after examining them in the Senckenbergian Museum, came to the correct conclusion that they belonged to a different species, on which he proposed to bestow the name Antilope ruficollis. We must therefore use Gazella ruficollis as the correct scientific designation of this animal. A third German naturalist, Heuglin, who has recorded his experiences of this species, tells us that he met with it, generally in pairs or small families, and often mixed up with herds of other Gazelles, in the desert districts of Dongola and Kordofan, where it is known to the Arabs as the “Adra” or “Ledra.” From this native name, Bennett, in his memoir on Gazella mhorr, to which we shall presently refer, proposed to call the present species Antilope addra, but, as has been already stated, Hamilton Smith’s name has precedence.

Gazella ruficollis is, we regret to say, represented in the British Museum by two specimens only, neither of which is suitable for exhibition in the[208] Gallery. One of these is a stuffed female from Sennaar, received from the Stockholm Museum, in exchange, in 1846, and the other an imperfect skin of a male from Kordofan, purchased of a dealer in 1848.

We are not aware that this Gazelle has ever been brought to Europe alive.

Our figure (Plate LXXI.) has been drawn from the stuffed female in the British Museum.

September, 1898.

[209]

105. THE DAMA GAZELLE.
GAZELLA DAMA (Pall.).

Size about as in G. ruficollis and G. mhorr. Markings throughout very much as in the next species, Gazella mhorr, but the white of the rump-patch, although less than in G. ruficollis, where it spreads all over the body, is considerably more extended, uniting on the thighs with the white of the sides of the belly, and therefore cutting off the dark colour of the outer sides of the hind limbs from that of the back. Other characters very much as in G. mhorr.

Passing now to the western end of Northern Africa we find this group of Antelopes represented by the “Dama” Gazelle, a species which has been known to naturalists ever since the time of Buffon. By him it was described and figured in the twelfth volume of his ‘Histoire Naturelle,’ under the name of “Le Nanguer” the appellation stated by Adanson to be given to it in Senegal. Upon the “Nanguer” of Buffon, Pallas in 1766 established his Antilope dama, so that there can be no question as to Gazella dama being the correct name of the representative species of this group in Senegal. But whether Pallas was right in assigning the term “dama” of Pliny to the present animal is a matter open to much question. The late Mr. E. T. Bennett has discussed this subject in his article on the Mhorr Antelope published in the first volume of the Zoological Society’s ‘Transactions,’ to which we may refer our readers. But there can be little doubt that the ordinary “Dama” of the Romans was not the present animal, but the well-known Fallow-Deer, Cervus dama.

For many years the naturalists following Buffon and Pallas gave us no further information concerning this Antelope, and merely copied what their predecessors had said of it. Sundevall, in his well-known treatise on the “Pecora,” united the three members of this group together under “Antilope[211] dama,” designating the forms of Morocco and Senegal as “var. occidentalis,” and stating that he had examined a specimen of it in the Frankfort Museum. If the locality and the references given by Rüppell in his catalogue of the Senckenbergian Collection are correct, the specimen in that collection must be the “Mhorr” of Morocco, and not the “Nanguer” of Senegal. We are told by Gray (‘Gleanings from the Knowsley Menagerie,’ p. 5) that the Frankfort specimen was originally received from the Zoological Society of London.

Among the numerous Antelopes procured from the Gambia for the Derby Menagerie by Lord Derby’s agent Whitfield were several examples of this species. Dr. Gray, in the letterpress to the ‘Gleanings,’ tells us that at the time of his writing (April 1850) there was a fine male of Gazella dama living at Knowsley, and that a female, procured by the same collector, which died on its passage home, was preserved in the British Museum. This specimen, a skin of an immature animal, is still in the Museum, to which it was presented by the Earl of Derby, and, strange to say, is still the only example of this Antelope in the National Collection.

In 1865 a pair of this Gazelle (as recorded in the Society’s ‘Proceedings,’ 1865, p. 675) were acquired by the Zoological Society of London, from the Zoological Gardens, Antwerp. About the same period Sclater recollects having seen other examples of this Antelope in the Antwerp Gardens, but does not remember to have noticed them in any other of the continental Gardens which he has visited from time to time for many years.

In some MS. notes on the Antelopes of the Gambia which Sir Robert Llewelyn, K.C.M.G., the Governor of the Colony, has kindly sent to Sclater, “the Springbuck,” or “Kongko-tong” of the Mandingos, is stated to be “common all over the south bank of the river.” This “Springbuck” can be hardly any other Antelope than the present species. If such is the case, it is remarkable that a beautiful animal, so abundant in a British Colony a few days’ steam from our shores, should be still so little known in our Museums and Menageries.

September, 1898.

[212]

THE BOOK OF ANTELOPES, PL. LXXII.

Wolf del. J. Smit lith.

Hanhart imp.

The Mhorr Gazelle.

GAZELLA MHORR.

Published by R. H. Porter.

[213]

106. THE MHORR GAZELLE.
GAZELLA MHORR (Benn.).
[PLATE LXXII.]

Size large, height of a male not fully adult 35½ inches. General colour dull rufous or rufous-fawn, becoming a deeper and richer rufous anteriorly. Face, cheeks, and chin whitish or whitish-fawn, the forehead beneath the horns and an inconspicuous dark cheek-streak blackish. Neck deep rufous, a conspicuous white spot on its front surface. No dark or light[214] lateral bands present, nor pygal bands. White of belly extending rather high up on the sides, the line of white passing across the outer sides of the forearms on to the chest, where it contrasts conspicuously with the dark red of the neck. White of rump much as in G. soemmerringi, including the tail, and extending angularly forwards on each side into the body-colour, but not uniting with the belly-colour across the outer sides of the thighs. All the boundary-lines between the rufous and white well defined. Knee-brushes small. Tail white, a small brownish or fawn-coloured tuft at its extremity.

Horns of male thick, strongly bent backwards below, recurved upwards and forwards above.

Female. Similar to the male, but horns shorter and thinner.

The last species of the group, and also of the genus Gazella, is the Mhorr, the Moroccan representative of the Dama Gazelle, to which, as we have already stated, it is very closely allied. Indeed, we have not inconsiderable doubts as to their real specific distinctness. Southern Morocco and Senegal approach so nearly together that it is not likely prima facie that the Gazelles of their deserts would be specifically different.

The well-known zoologist, Mr. E. T. Bennett, who was Secretary to the Zoological Society of London in its early days, was the first describer of the “M’horr” as distinct from Gazella dama, and brought his account of it before a Meeting of that Society on January 8th, 1833. His full memoir on this subject was afterwards published in the Society’s ‘Transactions,’ where it occupies the first pages of the first volume of that standard work.

Bennett’s observations were made on two specimens of this Gazelle which were presented, while living, to the Zoological Society by Mr. E. W. A. Drummond-Hay, C.M.Z.S., then British Consul-General at Tangier, for whom they had been procured by Mr. E. W. Willshire, C.M.Z.S., British Vice-Consul at Mogador. Bennett also had before him a third specimen in the shape of an imperfect skin of this Antelope, also presented to the Society by Mr. Drummond-Hay. All these specimens are stated to have been brought “from the territories of the Sheik of Wednoon, twelve days’ journey inland from the latter place”; but, in spite of the expression “inland,” we suppose “Ouednoon,” as it should be more correctly written, to be the valley[215] of the River Noon in the extreme southern coast district of Morocco, opposite the Canary Islands.

In the article in question, which is accompanied by an excellent coloured figure of the “M’hoor,” Bennett informs us that this Antelope “is regarded in the kingdom of Morocco as an exceedingly rare animal,” and continues as follows:—

About the same time as the Zoological Society’s specimens arrived in England it would appear that living examples of the same Antelope reached the Jardin des Plantes at Paris. One of these, an immature female, was figured by Geoffrey St.-Hilaire and F. Cuvier on the 375th plate of their ‘Histoire Naturelle des Mammifères.’ In the letterpress accompanying this plate we are informed that two young living examples (male and female) had been received there, and that the female had lived in good health for two years without changing her colour, only varying in the size of her body and in the shape of her horns. The authors inform us that when the “Nanguers” arrived at Paris they had only very short horns, about 4 or 5 inches in length, which were at that period strongly and uniformly curved towards the front. The male having died when young did not change the character of his horns, but those of the female having had time to develop became recurved behind and divergent one from the other. These Gazelles had neither tear-bags nor knee-brushes. When the figure was drawn the female stood about 2 feet 10 inches in height, but seemed to be not fully grown.

The authors of the ‘Histoire Naturelle des Mammifères’ refer these specimens to the “Nanguer” of Buffon, and do not say from what country they were received, but from their figure and description there can be no[216] doubt that they belonged to the Gazella mhorr of Morocco and not to the true G. dama of Senegal, if these two species are held to be distinct.

The only example of Gazella mhorr in the British Museum is the mounted adult male from Morocco presented to the Zoological Society by Mr. Willshire in 1833, and formerly in that Society’s collection. It is no doubt the specimen from which Bennett’s figure in the ‘Transactions’ was taken, and is also, we believe, the original of our illustration (Plate LXXII.), which was put upon the stone by Mr. Smit under the direction of the late Sir Victor Brooke. Further information concerning this and other animals of Southern Morocco is much required, but until the present political conditions of that country are altered we are hardly likely to obtain it.

September, 1898.

Size medium, about as in middle-sized Gazelles. Neck slender and rather elongated, but not so much as that of Lithocranius. Tail long, reaching nearly to the hocks. No Gazelline body-markings present, but the head with typically Gazelline streaks and bands. False hoofs small.

Skull long and low, approaching in shape that of Lithocranius, but less modified. Shallow anteorbital fossæ present. Premaxillæ reaching nasals. Premolars 3/3, the anterior in each jaw unusually small.

Horns of medium length, evenly divergent, curved upwards and forwards throughout their length, except just at their bases. Female hornless.

One species only.

THE BOOK OF ANTELOPES, PL. LXXIII.

J Smit del et lith

Hanhart imp.

The Dibatag.

AMMODORCAS CLARKEI.

Published by R. H Porter

[219]

107. THE DIBATAG.
AMMODORCAS CLARKEI (Thos.).
[PLATE LXXIII.]

Height at withers about 31 inches. General colour of body dark purplish rufous, no dark or light lateral or pygal bands present. Head with handsome sharply-defined Gazelline markings; the central facial band rich fulvous rufous, no darker spot on muzzle; light facial streaks long, sharply defined, and extending to the sides of the muzzle, pure white; areas round eyes and ears also whitish, though not sharply defined, connected with each other and continuous with the white facial streaks. Dark facial streaks present but not prominent. Back of ears dull whitish. Chin and interramia white. Cheeks below white markings, throat, and whole of neck dark purplish rufous, continuous with the back. Belly pure sharply-defined white, but on the chest this colour is broken by two projections running down from the dark throat-colour on each side of the middle line to between the elbows. Limbs, except their inner sides above, coloured like the back, but gradually passing lower down into fulvous instead of rufous. Knee-brushes present but small.[220] Back of hams white, the white not projecting into the dark body-colour. Tail long, well-haired, but not crested, blackish above and below, except just at its base.

Skull-dimensions in an adult male:—Basal length 7·5 inches, greatest breadth 3·7, muzzle to orbit 4·2.

Horns attaining about 11 or 12 inches in length; their basal halves broadly ringed, their long forwardly-directed terminal halves smooth and evidently corresponding to the short smooth recurved tips of ordinary Gazelline horns.

Female. Similar to the male, but without horns.

We now conclude the long series of Gazelles with three abnormal forms, each constituting a genus of itself, which, curiously enough, are all restricted to N.E. Africa. As regards the first two of them there can be no question, we believe, of their close alliance to the Gazelles, Ammodorcas being, in several respects, intermediate between Gazellla and Lithocranius, and leading on to that most specialized form of the group. About the correct position of the Beira, however, there is considerable doubt, and it is quite possible that a more natural place for it in the Antelopine series may be hereafter discovered.

The Dibatag, or Clarke’s Gazelle, was first met with by the sportsman and naturalist whose name it bears in 1890, during his exploring trip into the Dolbahanta and Marehán country south-east of Berbera. Writing to Thomas about his discovery, Mr. T. W. H. Clarke (of Secheron, Battery Point, Hobart, Tasmania) says (see P. Z. S. 1891, p. 209):—

Mr. Clarke’s skulls of this Antelope, which were at the time without head-skins, having been submitted to Thomas for examination by Messrs. Rowland Ward and Co. (to whom Mr. Clarke had sent for preservation his specimens of Antelopes obtained during this expedition), were, on account of the shape of the horns, not unnaturally supposed to belong to a new species of Reedbuck, and named “Cervicapra clarkei”—a mistake which Thomas corrected in his later paper. Very shortly after Mr. Clarke’s specimens had been sent to Europe Capt. Swayne purchased of an Arab in the market at Berbera two pairs of horns of this Antelope with the head-skins attached, and, seeing that they belonged to a new species, sent them to Sclater. Sclater exhibited these specimens at a meeting of the Zoological Society on March 17th, 1891, and pointed out their Gazelline affinities, but finding that they belonged to the same species as that just named by Thomas handed them over to the latter for further examination.

In his paper upon this subject, which was read at the same meeting of the Zoological Society, Thomas took the opportunity of describing the whole series of Mr. Clarke’s Antelopes, which were eight in number. For Clarke’s Gazelle, of which, with the aid of the head-skins and the cleaned skulls, he had no difficulty in recognizing the true affinities, he established the new[222] generic term “Ammodorcas,” and added a full description of this remarkable form, illustrated by two plates, one of which, by the kind permission of the Zoological Society, we are enabled to copy in the present work.

Fig. 83.

Head of the Dibatag, ♂.

(From P. Z. S. 1891, pl. xxi.)

As already pointed out, it is evident that in its skull-characters Ammodorcas is intermediate between Gazella and Lithocranius, while in the shape of its horns it is absolutely different from all other Gazelline genera.

[223]

Capt. Swayne, who has had the unequalled experience of seventeen visits to Somaliland, writing in 1894, says:—

Another well-known explorer of Somaliland, Mr. G. Percy V. Aylmer, kindly sends us the following field-notes on this species:—

Through the kind intervention of Mr. Pease we have also received some interesting particulars as to a wider range of the “Dibatag” from Lieut. R. Ward Jackson, of the 11th Hussars, who, accompanied by Capt. M. S. Wellby, has recently penetrated far into the eastern interior of the Somaliland peninsula. Mr. Jackson writes as follows:—

Fig. 84.

Map of Somaliland (showing the localities of the Dibatag).

[226]

The specimens of Ammodorcas clarkei in the British Museum consist of a head-skin with its skull from Burao Wells, presented by Mr. T. W. H. Clarke (the type of the species), and a stuffed male, the skins of a male and two females and their skulls from the same district, obtained in his second expedition, and presented by the same donor. There are also a skin and skeleton of a female from Darror Wells, on the Haud, presented by Mr. Ford G. Barclay, and the skull of a male from near the Bur Dap Mountain, Central Somaliland, presented by Col. A. Paget.

Our illustration of the male of this Gazelle, with the female in the distance (Plate LXXIII.), has been drawn and put on the stone from the specimens in the British Museum.

[Since this was written we have received from Dr. Matschie, through Herr Oscar Neumann, the interesting information that the Dibatag occurs in the Kilimanjaro district of German East Africa, a considerable extension of its previously known range.]

September, 1898.

Size about as in the larger Gazelles. Form very peculiar, owing to the great elongation of the neck, which gives almost a Giraffe-like appearance to the animal. Limbs also very long; false hoofs quite minute. Colouring of the head less typically Gazelline than in Ammodorcas, but a modification of the light lateral body-line present.

Skull (see fig. 85, p. 231) excessively long and low, the cranial part behind the horns particularly lengthened; the bones of this part very solid and stony. Bullæ low and opaque. A shallow anteorbital fossa present. Premaxillæ not reaching the nasals. Lower jaw slender and delicate. Premolars 3/3, the anterior one above almost as large as the second.

Horns thick, oval in section, very closely ringed, curved backwards as in many species of Gazella for the greater part of their length, their middle portion more or less lyrate, as in Gazella dorcas, their tips recurved upwards or forwards. Female hornless.

One species only.

THE BOOK OF ANTELOPES, PL. LXXIV.

J Smit del et lith.

Hanhart imp.

The Gerenuk.

LITHOCRANIUS WALLERI.

Published by R H Porter.

[229]

108. THE GERENUK.
LITHOCRANIUS WALLERI (Brooke).
[PLATE LXXIV.]

Height at withers 39 inches in an old male. General colour of back rich chestnut rufous, sharply bounded on the upperside of each flank by the very distinct light lateral line, which passes below imperceptibly into fawn-colour, much paler than the dorsal colour, no trace of a dark lateral line being present. The fawn-colour also extends on to the crown, all round the neck, on the shoulders, hips, and down the limbs. Central line of face deep rufous; area round eye whitish, sometimes extended forwards as a light final streak towards the muzzle, but more often stopping just anterior to the openings of[230] the lacrymal glands. Backs of ears pale fawn-colour; area round their bases but little lighter than rest of head. Fawn-colour of throat projecting down on to chest, as in Ammodorcas clarkei. Outer sides of limbs pale fawn throughout. Tail about 10 inches in length, rufous-fawn like the body basally above, whitish below, the tip tufted with black; on each side of its base the white of the hams, which is very narrow, runs up as a pointed projection into the dark body-colour.

The skull of an old male measures 8·75 inches in basal length, the greatest breadth being 3·8, and the muzzle to orbit 4·7.

The horns of good specimens attain 15 or 16 inches in length round the curves, but southern specimens, those from East Africa, have not, as a rule, such fine horns as those from Somaliland.

Female. Similar to the male, but without horns.

This Antelope was first made known to science by the late Sir Victor Brooke in 1878 in the last of the many excellent articles which he published in the Zoological Society’s ‘Proceedings.’ His description was based on two skulls with horns attached to them which were lent to him for examination by Mr. Gerald Waller, F.Z.S., at whose request they were named by Brooke after Mr. Waller’s brother, “who lost his life in Africa.”

Although it is not so stated by Brooke (who was presumably ignorant of the fact), Mr. Waller’s specimens of this Antelope are now known to have been given to him by Sir John Kirk, at that time H.B.M. Consul at Zanzibar. In reply to enquiries Sir John kindly informs us that these skulls were procured by hunters in his employment on the coast near the River Juba in Southern Somaliland. Sir John subsequently shot specimens of it himself in the same district (where at that period it was very common), and also brought living examples of it away to Zanzibar.

In his description Brooke pointed out that, as is well shown in the accompanying figure (p. 231), which has been copied from the plate that illustrates his paper, the skull of this Gazelle, besides its general depression, stands widely apart from those of all other species of the group in the enormous backward prolongation of the occiput—“an extension gained principally by the great size of the occipital bone and the prominence of the occipital crest.”[231] This divergence was so remarkable that Brooke doubted “whether the species should not constitute the type of a new subgenus.” As we shall see later, Brooke’s views have been fully justified by what has taken place since a close acquaintance with the structure of this singular Antelope has been acquired.

Fig. 85.

Skull of the Gerenuk.

(From Brooke, P. Z. S. 1878, pl. lvi.)

It was six years after the publication of Brooke’s paper before any additional information concerning this strange Antelope was obtained. In November 1884 Sclater brought before the notice of the Zoological Society a series of flat skins of Mammals, prepared by the natives of Somaliland, which had been lent to him for examination by Mr. C. Hagenbeck, of Hamburg. Amongst these were two skins, at first believed to belong to a new Gazelle, but which, after much research and mainly by the aid of a mounted head obtained in Somaliland by the late Mr. F. L. James, he was enabled to prove must belong to the same Antelope on the skull of which[232] Gazella walleri had been founded. The skins were at once distinguishable by the well-defined dark brown dorsal stripe which, as we now know, forms such a noticeable feature in the present species. Further evidence of the identity of the Somaliland Antelope with Gazella walleri was obtained by the comparison of Mr. James’s specimen with one of the typical skulls of the last-named species. They differed little, except in the slightly larger size of the northern specimen and in some other minor characters. It was thus first shown that the range of Waller’s Gazelle extends to Northern Somaliland, but we now know that the “Gerenuk,” as the Somalis call it, is one of the most abundant game-animals of that favoured land.

Fig. 86.

Sketch of Gerenuk, ♂ and ♀, in characteristic attitudes.

(From Neumann’s ‘Elephant-Hunting,’ p. 81. Lent by Messrs.
Rowland Ward & Co.)

Capt. Swayne sums up his long acquaintance with this Antelope as follows:—

Mr. Frederick Gillett, F.Z.S., who accompanied Dr. Donaldson Smith during the first part of his expedition to Lake Rudolph, has kindly drawn up for us the following notes on Waller’s Gazelle:—

We copy an account of the native mode of capture of this Gazelle from Messrs. Parkinson and Dunbar’s narrative of their journey in Northern Somaliland in 1896 (Geogr. Journ. xi. p. 25):—

Mr. Alfred Pease has kindly summarized his field-notes on the Gerenuk in the following paragraphs:—

We subjoin Mr. E. N. Buxton’s lively account of his experiences with this Antelope, extracted from the second series of ‘Short Stalks’:—

It will be recollected that Lithocranius walleri, although actually better known to us from Somaliland, was originally discovered in the southern part of its range, in British East Africa. Here, according to Mr. Hunter, it is “very rare in the Kilimanjaro district, though numerous up the Tana River.” Lieut v. Höhnel, who accompanied Count Teleki’s expedition to Lake Rudolph, informs us that it is common also on the Upper Tana, between Hameye and the mouth of the Mackenzie River, but that none were found on the Guaso Nyiro. Mr. Jackson, in the first volume of ‘Big Game Shooting,’ writes as follows concerning the habits of this species in British East Africa:—

The Gerenuk was well represented in the collection of Mammals obtained[237] by Mr. D. G. Elliot in the course of his recent expedition to Somaliland, and a series of specimens of this Antelope obtained on that occasion has been mounted in characteristic attitudes in one of the large show-cases of the Field-Columbian Museum at Chicago. Mr. Elliot has kindly sent us some photographs of this interesting group, which show the male, female, and young, and their peculiar manner of feeding.

The Gerenuk is represented in the British Museum by a fine mounted pair obtained by Herr Menges in the neighbourhood of Berbera, from which our illustration of both sexes (Plate LXXIV.) has been prepared by Mr. Smit. There are also four other skins of both sexes, purchased of Herr Menges, and the skin figured by Sclater (P. Z. S. 1884, pl. xlix.), which was presented to him by Herr Hagenbeck, but was likewise originally procured by Herr Menges. In the National Collection there are also a skin and skull of a male of this Antelope from Burao Wells, Somaliland, presented by Mr. T. W. H. Clarke, another skin and skull from the hills south of Berbera, presented by Capt. J. R. Harkness, R.A., and a mounted skeleton from Somaliland, obtained by purchase.

From the southern part of its range the British Museum has one of the original skulls from the River Juba district upon which the species was founded, and three skulls and some separate horns from the extreme south of the Somali coast, obtained by Sir John Kirk.

September, 1898.

Size very small, smaller than in any Gazelle. Ears large. Tail short. False hoofs minute; horny part of main hoofs short, but the internal pad to the hoof broad and thick; position of limbs in standing almost as in Oreotragus. No Gazelline head-markings, but a dark lateral band present on the sides of the body.

Skull broad and short. Nasals short and premaxillæ long, the structure in this region recalling that in Madoqua. Anteorbital region with a very small and shallow lacrymal fossa. Bullæ large and inflated. Premolars 3/3, the anterior very large.

Horns short, straight or faintly curved forwards, not unlike those of Raphiceros, and quite different from those of any true Gazelle. Female hornless.

One species only.

THE BOOK OF ANTELOPES, PL. LXXV.

Smit del. et lith.

Hanhart imp.

The Beira.

DORCOTRAGUS MEGALOTIS.

Published by R. H. Porter.

[241]

109. THE BEIRA.
DORCOTRAGUS MEGALOTIS (Menges).
[PLATE LXXV.]

Size about as in the Steinbok or Grysbok; height at withers 20 inches in an adult male. General colour of neck and back a peculiar purplish grey, very finely grizzled with white, the extreme tips of the hairs being this latter colour. Scarcely a trace of a light lateral band, but a distinct dark one present, brownish, passing quite across the shoulder, and extending behind on to the sides of the rump. Belly not, as is usual, pure white throughout, but only white on the axillæ and groins, the remainder pale yellowish fawn, or even orange-fawn, this colour also extending on to the outer sides of the forearms and thighs; rest of limbs becoming darker fulvous to the hoofs. Head bright fulvous, quite different to the neck; area round eyes white, but no Gazelline facial streaks present. Tail short and rather bushy, coloured like the back, without darker tip.

Measurements of an old male skull:—Basal length 5·35 inches, greatest breadth 3·15, muzzle to orbit 3·2.

Horns ordinarily from 4 to 5 inches in length, the longest in the British Museum collection, one of Capt. Swayne’s, attaining 5½ inches.

[242]

On several occasions during the many years in which Sclater’s excellent correspondent Captain H. G. C. Swayne, R.E., was engaged on his various explorations and expeditions in Somaliland, he wrote about a “small red Antelope” found in the mountains which had the habits of a “Klipspringer,” but, according to native testimony, was of quite a different species. The existence of the same animal had also been recorded as long ago as 1885 under the name “Behra,” by Herr Josef Menges, in an account of his fourth journey in Somaliland, published in Petermann’s ‘Mittheilungen.’ Herr Menges met with the “Behra” on the Hekebo plateau (about 10° S. lat. and 44° 40´ E. long.), and had at one time a young living specimen of it in his possession.

After Herr Menges the “Behra” or “Beira,” as it is now usually called, after its Somali name, seems to have been first actually seen by Lieut. E. J. Swayne, of the Indian Staff Corps, Capt. Swayne’s brother, when he was in the Gadabursi country in the autumn of 1891. He observed two of them among very rugged hills, but failed to get a shot at them. He described them to Capt. Swayne as being “reddish Antelopes, rather larger than the Klipspringer, with small straight horns, which bounded away among the rocks in exactly the same manner as the Klipspringer.”

Capt. Swayne was much excited about this discovery, and promised Sclater to do all he could to procure specimens of the animal. On his last trip to Somaliland he was assured by his Somalis that he would find the “Beira” on Waggar Mountain, near the south-eastern extremity of the Golis range, but he had not time to go there. On leaving Berbera, however, Capt. Swayne exhorted his men to proceed to the mountains themselves and to endeavour to procure some specimens of the Beira, offering them a handsome reward for good heads and skulls of a male and female, and leaving instructions to his agents there to pay the men and to forward the specimens.

Early in 1894 the much-wished-for skins were obtained by the faithful Somalis and forwarded to Sclater by Captain Swayne. We were proceeding to describe and figure them in the Zoological Society’s ‘Proceedings,’ when we found that we had been anticipated by Herr Menges, who had just[243] described the species in the ‘Zoologischer Anzeiger,’ and given it the name “Oreotragus megalotis.” It has, however, certainly but a very remote connection, except as regards its habits, with the Klipspringer (Oreotragus), and Dr. Noack was quite justified in proposing for it the new and appropriate term Dorcatragus (δορκας, an Antelope, and τραγος, a Goat), which he did in the same periodical shortly afterwards. Dr. Noack based his article upon two specimens, male and female, which he had then lately received from Herr Menges for examination.

The next traveller in Somaliland to encounter this rare Antelope was, we believe, Capt. P. Z. Cox, who, writing to Dr. Günther from Berbera in April 1895, gave an account of the circumstances under which he obtained a female example of the Beira for the British Museum:—

Sir Edmund Loder has kindly favoured us with the following field-notes on the Beira:—

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The accompanying view of the head is enlarged from a photograph of this Antelope kindly sent to us by Sir Edmund Loder.

Fig. 87.

Front view of the head of the Beira.

During Mr. D. G. Elliot’s East-African expedition of 1896, Mr. Akeley, one of his party, obtained two examples of this Antelope on Nasr Hablod Mountain, near Hargeisa, in nearly the same district as that just described by Sir Edmund Loder. These specimens are now in the Field-Columbian Museum at Chicago.

The British Museum contains a mounted specimen of an adult male of the Beira (from which our figure, Plate LXXV., has been prepared by Mr. Smit) and a skin of a female of the same species purchased of Herr Menges, besides the two skins obtained by the Somalis for Capt. Swayne, as above mentioned, which have been presented to the Museum by Sclater. It contains also a skin and complete skeleton of a female of this Antelope presented by Capt. P. Z. Cox, as described above.

September, 1898.

Transcriber’s Notes:

1. Obvious printers’, punctuation and spelling errors have been corrected silently.

2. Some hyphenated and non-hyphenated versions of the same words have been retained as in the original.

3. Where hyphenation is in doubt, it has been retained as in the original.